| Mary E. Barkworth |
1 |
Lower glumes
of the sessile spikelets not winged, rugose, with 4-5 ridges; plants annuals
or short-lived perennials ..... 1. I.
rugosum |
Lower glumes of the sessile spikelets
winged on the keels, not rugose; plants perennial ..... 2. I.
indicum |
1. Ischaemum rugosum Salisb.
Ribbed Murainagrass
Plants annual, or short-lived perennials. Culms 30-130 cm, erect
or geniculate at the base, simple to strongly branched below; nodes antrorsely
pilose; internodes glabrous. Leaves cauline; sheaths mostly
glabrous or sparsely pilose, margins ciliate distally; collars villous;
ligules 1-5.5 mm; blades 8-20 cm long, 7-15 mm wide, flat, usually
pilose on both surfaces, sometimes with papillose-based hairs, occasionally glabrous.
Inflorescence units with 2 rames; rames 3-8 cm long, 3-4 mm wide;
internodes 2.5-3.5 mm, slightly clavate distally. Sessile spikelets
3-5 mm long, 1.8-2.2 mm wide; calluses shortly pubescent; lower glumes
coriaceous, yellowish, not winged, coarsely transversely rugose with 4-5 ridges
on the proximal 2/3-4/5, thickly chartaceous distally and tapering to the apices;
upper glumes thickly chartaceous, ciliate, awned, awns 1.5-2 cm, geniculate
below the middle; anthers about 2 mm. Pedicellate spikelets varying
from 0.5 mm to equaling the sessile spikelets. 2n = 18, 20, 44.
Ischaemum rugosum is native to southern Asia, and is now established in
moist, tropical habitats around the world, including Mexico. It has been found
in southern Texas and on chrome ore piles in Canton, Maryland, but is thought
to have been eliminated from both areas. The U.S. Department of Agriculture considers
it a noxious weed; plants found growing within the continental United States should
be promptly reported to that agency.
2. Ischaemum indicum (Houtt.) Merr.
Indian Murainagrass
Plants perennial, forming loose clumps. Culms 20-90 cm, decumbent,
often rooting from the lower nodes; nodes antrorsely pilose; internodes
glabrous. Leaves crowded towards the base; sheaths open,
sparsely to densely pilose; ligules about 1 mm, membranous; blades 4-15
cm long, 3-8 mm wide, pilose on both surfaces. Inflorescence units with
2 rames; rames 3-5 cm long, 4-5 mm wide; internodes 3-4
mm long, 0.5-0.7 mm wide, slightly triquetrous. Sessile spikelets 4-5
mm long, 1.2-1.5 mm wide; calluses strigose; lower glumes coriaceous
below, thickly membranous above, rough but not rugose, 5-7-veined, 2-keeled,
keels narrowly winged;
upper glumes 1-keeled, keels winged distally, awned, awns 1-1.5
cm; anthers
2-2.5 mm. Pedicels 2.5-3.5 mm, ciliate on the edges. Pedicellate
spikelets
slightly smaller than the sessile spikelets. 2n = 36.
Reed (1964) reported finding Ischaemum indicum growing on chrome ore piles
in Canton, Maryland. This reported introduction has not been verified.