| John W. Thieret† |
Plants annual or perennial; scrambling.
Culms 0.5-2 m, ascending to decumbent, often rooting at the nodes, branched.
Leaves not aromatic; sheaths open, at least the outer margins
pubescent, usually with papillose-based hairs; ligules membranous, fimbriate
or ciliate; blades ovate to ovate-lanceolate. Inflorescences terminal
and axillary, panicles of subdigitate, often flabellate, clusters of rames;
rame internodes not sulcate; disarticulation in the rames, beneath
the sessile spikelets. Spikelets in heteromorphic sessile-pedicellate
pairs or appearing solitary and sessile, pedicels greatly reduced and lacking
spikelets. Sessile spikelets bisexual, with 2 florets; calluses
absent or blunt; glumes equal or subequal; lower florets sterile,
reduced to an unawned lemma; upper florets bisexual, awned (rarely unawned);
anthers 2 or 3. Pedicels 0.2-3 mm, not thickened, not fused to
the rame axes. Pedicellate spikelets absent or rudimentary. x
= 9, 10. Name from the Greek arthron, segment, and axon, axis,
referring to the jointed inflorescence axes.
Arthraxon is a genus of seven species that are native to tropical and subtropical
regions of the Eastern Hemisphere; one species is established in the Flora
region.
1. Arthraxon hispidus (Thunb.) Makino
Jointhead, Small Carpetgrass
Plants annual. Culms 0.5-1(2) m, weak, often decumbent and rooting
at the lower nodes; nodes hispid. Leaves cauline; sheaths
usually shorter than the internodes; ligules 0.4-3.5 mm, ciliate; lower
blades 1-7.5 cm long, 4-20 mm wide, cordate-clasping, ovate to ovate-lanceolate,
flat, margins ciliate (sometimes sparingly so), surfaces usually glabrous, abaxial
surface rarely hispidulous; upper blades greatly reduced. Panicles
1.3-7 cm, flabellate or contracted, with 12-20 rames; rames 1-6(11) cm.
Sessile spikelets: glumes 3-5.5 mm, lanceolate; lower glumes
several-veined; upper glumes 1- or 3-veined; awns 0.3-9 mm, included
or exserted, usually twisted below, sometimes geniculate at midlength; anthers
usually 2, 0.5-0.7 mm. Pedicels absent or to 2 mm. Pedicellate spikelets
absent. 2n = 36.
Arthraxon hispidus is native to Asia, but is naturalized and spreading
along roadsides, shores, ditches and in low woods and fields of the eastern
United States. It is also naturalized in Mexico, Central America, and
the West Indies. Plants in the Flora region belong to A. hispidus (Thunb.)
Makino var. hispidus, the most widespread and variable of the four
varieties.
Arthraxon castratus (Griff.) V. Naray. ex Bor, reported from
Puerto Rico, differs from A. hispidus in having pilose lemma margins,
a palea in its second floret, and three anthers.