| Christopher S. Campbell |
Plants perennial; usually cespitose,
sometimes rhizomatous. Culms 20-310 cm, erect, much-branched distally.
Leaves not aromatic; ligules membranous, sometimes ciliate; blades
linear, flat, folded, or convolute. Inflorescences terminal and axillary
or a false panicle; inflorescence units 1-600+ per culm; peduncles
initially concealed by the subtending leaf sheaths, sometimes exserted beyond
the sheaths at maturity, with (1)2-5(13) rames; rames not reflexed at
maturity, axes slender, terete to flattened, not longitudinally grooved, usually
conspicuously pubescent, with spikelets in heterogamous sessile-pedicellate
pairs (the terminal spikelets sometimes in triplets of 1 sessile and 2 pedicellate
spikelets), apices of the internodes neither cupulate nor fimbriate; disarticulation
in the rames, below the sessile spikelets. Sessile spikelets bisexual,
awned, with short, blunt calluses; lower glumes 2-keeled, flat or concave,
usually not veined between the keels, sometimes 2-9-veined; anthers 1,
3(2). Pedicels usually longer than 3 mm, similar to the rame internodes
in shape, length, and pubescence color, not fused to the rame axes. Pedicellate
spikelets usually vestigial or absent, sometimes well-developed and staminate.
x = 10. Name from the Greek andro, man, and pogon, beard,
referring to the pubescent pedicels of the staminate spikelets.
Andropogon is a cosmopolitan genus of tropical and temperate zones, comprising
approximately 120 species. Thirteen species are native to the Flora region.
Andropogon bicornis has been found
in the region, but is not known to be established. All but A.
hallii grow in the southeastern United States.
Several taxa are ecologically important in North America. Andropogon
gerardii is one of the most important native grasses in North America,
being one of the dominant species in the tallgrass prairies that used to cover
the center of the continent. Many varieties of A.
glomeratus and A. virginicus
aggressively colonize abandoned fields, cutover timberlands, and roadsides.
Some species are used in restoration and landscaping.
Species of Andropogon with solitary rames are easily confused with Schizachyrium
but, in Andropogon, the lower glumes of the sessile spikelets are flat
or concave and the rame internodes are not cupulate, whereas Schizachyrium
has convex glumes and rame internodes with strongly cupulate apices. Successful
identification of species in Andropogon
sect. Leptopogon (numbers 3-14) requires mature, complete specimens
and careful field study (Campbell 1983, 1986).
1 |
Pedicellate
spikelets usually well-developed, (3.5)6-12 mm long, usually staminate;
sessile spikelets 5-12 mm long (sect.
Andropogon) (2) |
Pedicellate spikelets usually vestigial
or absent, those of the terminal spikelet units occasionally well-developed
and staminate; sessile spikelets 2.6-8.4 mm long (sect.
Leptopogon) (3) |
|
Sessile spikelets with
awns 8-25 mm long; ligules 0.4-2.5 mm long; hairs of the rame internodes
2.2-4.2 mm long, sparse to dense; rhizomes sometimes present, the internodes
usually less than 2 cm .... 1. A.
gerardii |
|
Sessile spikelets unawned or with
awns less than 11 mm long; ligules (0.9)2.5-4.5 mm long; hairs of the
rame internodes 3.7-6.6 mm long, usually dense; rhizomes always present,
the internodes often more than 2 cm long ..... 2. A.
hallii |
|
Peduncles with solitary
rames; plants of southern Florida ..... 3. A.
gracilis |
|
Peduncles with (1)2-13 rames; plants
of varied distribution, including southern Florida (4) |
|
Rames not or scarcely
exserted at maturity; peduncles mostly less than 15 mm long at maturity
(5) |
|
Rames sometimes exserted above
their subtending sheaths at maturity; 1 or more peduncles more than 15
mm long at maturity (8) |
|
Culms 30-140 (usually
about 80) cm tall; blades 0.8-5 (usually about 2.5) mm wide; inflorescence
units 2-31 per culm ..... 9. A.
gyrans |
|
Culms 20-250 (usually more than
90) cm tall; blades 1.7-9.5 (usually more than 3) mm wide; inflorescence
units 3-600 per culm (6) |
|
Blades pubescent, most
hairs appressed; callus hairs 1.5-5 mm long ..... 13. A.
longiberbis |
|
Blades glabrous or with spreading
(rarely appressed) hairs; callus hairs 1-3 mm long (7) |
|
Blades 11-52 cm long;
sheaths smooth, rarely somewhat scabrous; ligules 0.2-1 mm long; keels
of the lower glumes usually smooth below midlength, scabrous distally .....
12. A. virginicus |
|
Blades 13-109 cm long; sheaths
usually scabrous; ligules 0.6-2.2 mm long; keels of the lower glumes sometimes
scabrous below midlength ..... 14. A.
glomeratus |
|
Anthers 3 (9) |
|
Anthers 1 (rarely 3) (10)
|
|
Sessile spikelets 4.5-8.4
mm long; pedicellate spikelets 1.5-3.6 mm long, sterile; plants common
and widespread in the southeastern United States ..... 4. A.
ternarius |
|
Sessile spikelets 3-4 mm long;
pedicellate spikelets mostly vestigial or absent, those of the terminal
spikelet units well-developed and staminate; in the Flora region,
known only from southern Florida ..... 5. A.
bicornis |
|
Peduncles
with 2-13 rames ..... 8. A. liebmannii |
|
Peduncles usually with 2 (infrequently
up to 4) rames or (in A. gyrans var. gyrans and A. virginicus
var. virginicus), 2-5 (infrequently up to 7) rames (11) |
|
Culms 30-120(140) (usually
less than 100) cm tall; blades 0.8-5 (usually less than 3) mm wide; inflorescence
units 2-31 per culm (12) |
|
Culms (20)90-310 (usually more
than 100) cm tall; blades 1.7-9.5 (usually more than 3) mm wide; inflorescence
units 5-210 per culm (13) |
|
Peduncles with 2-5 rames;
anthers 0.6-1.7 mm long; sessile spikelets (3)4.1-4.4(5.7) mm long ..... 9.
A. gyrans |
|
Peduncles with 2 rames; anthers
1.2-2 mm long; sessile spikelets (4)4.8-5(5.5) mm long ..... 10. A.
tracyi |
|
Upper portion of the
plants open, the branches conspicuously arching ..... 11. A.
brachystachyus |
|
Upper portion of the plants dense,
the branches usually straight and erect to ascending (14) |
|
Rame internodes usually
densely and uniformly pubescent over their entire length; anthers 1.3-3.5
mm long; sessile spikelets (3.8)4-6.1 mm long (15) |
|
Rame internodes sparsely pubescent
basally, more densely pubescent distally; anthers 0.5-1.5 mm long; sessile
spikelets 2.6-4(5) (16) |
|
Blades 15-35 cm long,
often more or less pubescent; sheaths smooth, very rarely somewhat scabrous;
anthers 2-3.5 mm long; inflorescence units 5-45 per culm ..... 6. A.
arctatus |
|
Blades 32-61 cm long, usually glabrous;
sheaths often scabrous; anthers 1.3-2 mm long; inflorescence units usually
at least 50 (9-210) per culm ..... 7. A. floridanus |
|
Blades 11-52 cm long;
sheaths smooth, rarely somewhat scabrous; ligules 0.2-1 mm long; keels
of the lower glumes usually smooth below midlength, scabrous distally
..... 12. A. virginicus |
|
Blades 13-109 cm long; sheaths
usually scabrous, sometimes smooth; ligules 0.6-2.2 mm long; keels of
the lower glumes sometimes scabrous below midlength ..... 14. A.
glomeratus |
Andropogon L. sect. Andropogon
Inflorescences usually only terminal, axillary inflorescences
absent or few. Lower glumes thinly coriaceous, with 3-9 veins between
the keels; keels often winged. Pedicellate spikelets usually well-developed,
staminate.
1. Andropogon gerardii Vitman
Big Bluestem, Barbon de Gerard
Plants often forming large clumps, rhizomes, if present,
with internodes shorter than 2 cm. Culms 1-3 m, often glaucous. Sheaths
glabrous or pilose; ligules 0.4-2.5 mm; blades 5-50 cm long,
(2)5-10 mm wide, usually pilose adaxially, at least near the collar. Inflorescence
units usually only terminal; peduncles with 2-6(10) rames; rames
5-11 cm, exserted at maturity, usually purplish, sometimes yellowish; internodes
sparsely to densely pubescent, hairs 2.2-4.2 mm, usually white, rarely
yellowish.
Sessile spikelets 5-11 mm, scabrous; awns 8-25
mm; anthers 3, 2.5-4.5 mm. Pedicellate spikelets 3.5-12
mm, usually well-developed and staminate. 2n = 20, 40, 60 (usually),
70, 80, 90.
Andropogon gerardii grows in prairies, meadows, and generally dry soils.
It is a widespread species, extending from southern Canada to Mexico, and was
once dominant over much of its range. It is frequently planted for erosion control,
restoration, or as an ornamental; the records from Washington and central Montana
reflect such plantings. It hybridizes with A.
hallii, the two sometimes being treated as conspecific subspecies.
2. Andropogon hallii Hack.
Sand Bluestem
Plants strongly rhizomatous, rhizome internodes often longer than 2 cm.
Culms (40)60-150(200) cm, strongly glaucous. Ligules (0.9)2.5-4.5
mm, ciliate; blades 3-40(51) cm long, (1.5)2-10 mm wide, often pilose,
at least near the collar. Inflorescence units usually only terminal;
peduncles with 2-7 rames; rames 4-7(9) cm, exserted at maturity;
internodes usually densely pubescent, hairs 3.7-6.6 mm, often strongly
yellowish. Sessile spikelets (5)6.5-12 mm; lower glumes often
ciliate; awns absent or to 11 mm; anthers 3, (2.3)4-6 mm. Pedicellate
spikelets 3.5-12 mm, usually well-developed and staminate. 2n = 60
(usually), 70, 100.
Andropogon hallii grows on sandhills and in sandy soil. Its range extends
through the central plains into northern Mexico. It is similar to A.
gerardii, differing primarily in its rhizomatous habit, more densely
pubescent rames and pedicels, and greater drought tolerance. Andropogon hallii
and A. gerardii are sympatric in some locations. The two species can
hybridize and are sometimes treated as conspecific subspecies.
Andropogon sect. Leptopogon Stapf
Inflorescences false panicles; inflorescence units
numerous. Lower glumes membranous; keels not winged, intercostal
region usually not veined or with inconspicuous veins. Pedicellate spikelets
usually vestigial or absent.
3. Andropogon gracilis Spreng.
Wire Bluestem
Plants densely cespitose. Culms 20-60 cm, wiry, glabrous. Sheaths
smooth; ligules to 1.4 mm; blades to 45 cm long, to 4 mm wide, involute
and filiform, or folded. Inflorescence units 3-50+ per culm; peduncles
2-13.2 cm, with 1 rame; rames 2-4 cm, usually long-exserted at maturity;
internodes densely pubescent, hairs to 8 mm. Sessile spikelets 4-6
mm; lower glumes scabrous in the distal 1/2; awns 11-20 mm. Pedicellate
spikelets reduced to an awned or unawned glume, sterile. 2n = 40.
Andropogon gracilis grows on oölite in openings and rocky margins
of pine woodlands of southern Florida and the West Indies. Although not uncommon,
it is frequently overlooked. It has sometimes been placed in Schizachyrium because of its solitary rames.
4. Andropogon ternarius Michx.
Split Bluestem
Plants cespitose. Culms 70-150 cm. Sheaths smooth or scabrous,
sometimes pilose; ligules 0.4-1.5 mm, ciliate; blades 1-3 mm wide,
pubescent or glabrous and glaucous. Inflorescence units 2-30+ per culm;
peduncles usually 5-20 mm, with (1)2 rames; rames 3-4 cm, exserted
at maturity, terminating in a sessile-pedicellate spikelet pair; internodes
sparsely to densely villous, hairs from as long as to twice as long as the sessile
spikelets. Sessile spikelets 4.5-8.4 mm; callus hairs to 8 mm; awns
10-25 mm; anthers 3, 1.2-2.3 mm. Pedicellate spikelets 1.5-3.6
mm, sterile. 2n = 40, 60.
Andropogon ternarius grows in the southeastern United States and northern
Mexico. It is planted as an ornamental and for erosion control on slopes in
poor and sandy soils, and is tolerant of coastal conditions.
Andropogon ternarius is similar to A.
arctatus, but differs in its possession of three anthers and usually
in its longer spikelets, both sessile and pedicellate.
1 |
Rames densely villous, with
hairs about twice as long as the sessile spikelets and more or less obscuring
them; lower glumes of the sessile spikelets sometimes scabrous, without
conspicuous veins between the keels ..... var. cabanisii |
Rames sparsely villous, with hairs about as long as
the sessile spikelets, but not obscuring them; lower glumes of the sessile
spikelets scabrous, often conspicuously 2-veined between the keels .....
var. ternarius |
Andropogon ternarius var. cabanisii (Hack.) Fernald
& Griscom
Rames densely villous; internode hairs about twice as long as and
more or less obscuring the sessile spikelets. Lower glumes of sessile spikelets
glabrous, sometimes scabrous, not conspicuously veined between the keels.
Andropogon ternarius var. cabanisii grows in dry pine woods and
scrublands of peninsular Florida.
Andropogon ternarius Michx. var. ternarius
Rames sparsely villous; internode hairs about as long as, but not
obscuring, the sessile spikelets. Lower glumes of sessile spikelets scabrous,
often with 2 conspicuous veins between the keels.
Andropogon ternarius var. ternarius grows in dry, sandy woods, fields,
openings, and roadsides of the southeastern United States and Mexico.
5. Andropogon bicornis L.
Barbas de Indio
Plants densely cespitose, upper portion dense, obovate
to obpyramidal. Culms60-250 cm; internodes not glaucous. Sheaths
smooth; ligules 0.6-1 mm; blades 20-70 cm long, 2-7 mm wide, usually
glabrous or scabrous on the margins. Inflorescence units 50-500; subtending
sheaths 2.5-4.5 cm long, 2-3 mm wide; peduncles 20-70 mm, with 2(3)
rames; rames 2-4 cm, exserted at maturity; internodes filiform,
densely and evenly pubescent, hairs 3-9 mm. Sessile spikelets 3-4 mm;
unawned; callus hairs 0.5-1 mm; keels of lower glumes scabrous
above the midpoint; anthers 3, 1-1.4 mm. Pedicellate spikelets
mostly vestigial or absent, 1-2 of those in the terminal units on each rame
3-5 mm and staminate. 2n = 60, 120.
Andropogon bicornis is a widespread species of the Western Hemisphere
tropics. It was collected in the early 1960s in Dade County, Florida, near the
track of a major hurricane, but may not be established in the Flora region.
6. Andropogon arctatus Chapm.
Pinewoods Bluestem
Plants cespitose or somewhat rhizomatous, upper portion dense, oblong to
ovate. Culms 90-170 cm; internodes occasionally somewhat glaucous
just below the nodes; branches straight, erect to ascending. Sheaths
smooth, rarely somewhat scabrous; ligules 0.3-0.9 mm, sometimes ciliate,
cilia to 0.5 mm; blades 15-35 cm long, 3-8 mm wide, glabrous or densely
pubescent, hairs spreading. Inflorescence units 5-45 per culm; subtending
sheaths (3.3)3.8-6.8(9) cm long, (2.5)3.2-4(5) mm wide; peduncles (9)26-66(115)
mm, with 2(4) rames; rames (2.2)2.6-4.3(5.3) cm, usually exserted at maturity,
pubescence either evenly distributed or more dense distally within each internode.
Sessile spikelets (4.3)4.9-5.4(6.1) mm; callus hairs 1.5-2.5 mm;
keels of lower glumes scabrous from below the midpoint; awns 5-16
mm; anthers 1(3), 2-3.5 mm, red. Pedicellate spikelets vestigial
or absent. 2n = 20.
Andropogon arctatus grows in flatwoods, bogs, and scrublands of southern
Alabama and Florida. Its flowering appears to be stimulated by fire but, unlike
other members of sect. Leptopogon in the
Flora region, the effect lasts only one or two years, the plants then remaining
vegetative until the next fire occurs. It is similar to A.
ternarius, but differs in its long, usually solitary anther and shorter
spikelets.
7. Andropogon floridanus Scribn.
Florida Bluestem
Plants cespitose, usually densely obpyramidal to oblanceolate above. Culms
70-210 cm; internodes occasionally somewhat glaucous just below the node;
branches straight, mostly erect to ascending. Sheaths often scabrous,
sometimes smooth; ligules 0.4-1.2 mm, ciliate, cilia 0.2-1.3 mm; blades
32-61 cm long, 2.9-5 mm wide, glabrous, rarely sparsely pubescent. Inflorescence
units (9)50-210 per culm; subtending sheaths (3)4-5.9(7) cm long, (1.5)2-2.7(3.6)
mm wide; peduncles (10)19-48(93) mm, with 2(4) rames; rames (2)2.5-3.7(4.5)
cm, usually exserted at maturity, internodes evenly pubescent. Sessile
spikelets (3.8)4.4-4.8(5.5) mm; callus hairs 1-3 mm; keels of lower
glumes glabrous below midlength; awns 5-15 mm; anthers 1(3),
1.3-2 mm, usually yellow (sometimes purple). Pedicellate spikelets vestigial
or absent. 2n = 20.
Andropogon floridanus grows on sandy soils in southeastern Georgia and
Florida, being most abundant in Pinus clausa scrublands. It usually occurs
in small stands, but stands of about a hundred individuals have been observed.
8. Andropogon liebmannii Hack.
Liebmann's Bluestem, Mohr's Bluestem
Plants cespitose, cylindrical to oblong above. Culms 20-170 cm;
internodes not glaucous; branches mostly erect, straight. Sheaths
smooth; ligules 0.7-1.2 mm, sometimes ciliate, cilia to 0.4 mm; blades
3-35 cm long, 2.5-7.5 mm wide, sparsely to densely pubescent with spreading,
shaggy hairs. Inflorescence units 7-50 per culm; subtending sheaths
(4)4.9-7.4(10) cm long, (3)4.2-6.1(10.1) mm wide; peduncles (10)24-68(130)
mm, at least some extending beyond the subtending sheaths at maturity, with
2-13 rames; rames (2)2.4-4(5) cm, usually exserted at maturity, pubescence
increasing in density distally within each internode. Sessile spikelets
(3)4-4.5(6.9) mm; keels of lower glumes scabrous above (and sometimes
below) the midpoint; awns 17-24 mm; anthers 1, 0.7-1.4 mm, yellow.
Pedicellate spikelets vestigial or absent. 2n = 20.
Andropogon liebmannii has two varieties. Andropogon liebmannii
var. pungensis (Ashe) C.S. Campb., the variety found in the Flora
region, differs from A. liebmannii Hack. var. liebmannii, which
grows in Mexico, in having culms that are usually more than 80 cm tall, leaves
that are more than 15 cm long, and sessile spikelets that are more than 4.2
mm long; in var. pungensis the culms are usually less than 90 cm tall,
the leaves less than 15 cm long, and the sessile spikelets less than 4.2 mm
long.
Andropogon liebmannii var. pungensis grows along
the coastal plain of the southeastern United states in bog, swamp, savannahs,
and flatwoods. It used to be treated as a species, which was known as A.
mohrii (Vasey) Hack. The English name 'Mohr's Bluestem' reflects this treatment.
9. Andropogon gyrans Ashe
Plants cespitose, cylindrical to ovate above. Culms 30-100(140)
cm; internodes usually glaucous; branches mostly erect, straight.
Sheaths smooth; ligules 0.3-1.5 mm, sometimes ciliate, cilia to
0.7 mm; blades 6-48 cm long, 0.8-5 mm wide, glabrous or densely pubescent
with spreading hairs. Inflorescence units 2-31 per culm; subtending
sheaths (2.6)4.1-4.5(13.5) cm long, (1.5)2.7-4.7(8) mm wide; peduncles
(1)5-31(195) mm, with 2-5 rames; rames (1.5)2.8-4.2(6) cm, exserted or
not at maturity, pubescence increasing in density distally within each internode.
Sessile spikelets (3)3.9-4.7(5.7) mm; callus hairs 1-5 mm; keels
of lower glumes scabrous only beyond midlength; awns 8-24 mm; anthers
1, 0.6-1.4(1.7) mm, yellow or purple. Pedicellate spikelets vestigial or
absent. 2n = 20.
Andropogon gyrans extends from the southeastern United States to the Caribbean
and Central America.
1 |
Ligules 0.3-1.1 mm long; rames usually hidden
within the more or less overlapping and inflated upper sheaths at maturity;
plants usually of well-drained soils ..... var. gyrans |
Ligules 0.8-1.5 mm long; rames usually exposed at maturity;
plants of wet habitats ..... var. stenophyllus |
Andropogon gyrans Ashe var. gyrans
Elliott's Beardgrass
Ligules 0.3-0.8(1.1) mm. Inflorescence units usually with 2-5
rames; rames usually concealed at maturity.
Andropogon gyrans var. gyrans generally grows in dry, sandy soil
of roadsides, embankments, fields, and pine or oak woods, occasionally in moister
soil. Its range extends south from the United States to the Caribbean and Central
America. Plants from Florida and Mississippi do not have inflated sheaths.
Andropogon gyrans var. stenophyllus (Hack.) C.S.
Campb.
Ligules (0.8)1.1-1.5 mm. Inflorescence units usually with 2 rames;
rames usually exposed at maturity.
Andropogon gyrans var. stenophyllus grows in ditches, bogs, savannahs,
and pond margins of the coastal plain, from eastern Texas to North Carolina.
10. Andropogon tracyi Nash
Tracy's Bluestem
Plants cespitose, upper portion dense, cylindrical. Culms 50-120
cm; internodes not glaucous; branches mostly erect, straight. Sheaths
smooth; ligules 0.2-0.5 mm, ciliate, cilia 0.2-0.8 mm; blades 10-22
cm long, 1.2-2.6 mm wide, glabrous or sparsely pubescent, with spreading hairs.
Inflorescence units 3-11 per culm; subtending sheaths (2.8)4.1-5.8(7.2)
cm long, (3)4-4.7(5.8) mm wide; peduncles (9)14-31(65) mm, with 2 rames;
rames (1.5)2.4-3.6(4.2) cm, usually exserted at maturity, pubescence increasing
in density distally within each internode. Sessile spikelets (4)4.8-5(5.5)
mm; callus hairs 1.5-3.5 mm; keels of lower glumes scabrous only
above the midpoint; awns 11-23 mm; anthers 1, 1.2-2 mm, yellow.
Pedicellate spikelets vestigial or absent. 2n = 20.
Andropogon tracyi grows on sandhills, sandy pinelands, and scrublands of
the southeastern United States. It resembles A. longiberbis,
but usually differs in having sparsely pubescent blades and a more slender appearance.
11. Andropogon brachystachyus Chapm.
Shortspike Bluestem
Plants cespitose, open and ovate to obpyramidal above. Culms 1.1-3.1
m; internodes not glaucous; branches arching. Sheaths smooth;
ligules 0.2-0.5 mm, ciliate, cilia 0.6-1.5 mm; blades 21-54 cm long,
2.3-6 mm wide, glabrous or sparsely pubescent, with spreading hairs. Inflorescence
units 12-190 per culm; subtending sheaths (2.1)2.4-3.5(4.1) cm long,
(2.3)2.6-3(3.8) mm wide; peduncles (13)20-31(43) mm, with 2(3) rames; rames
(1.2)1.5-2.1(2.6) cm, usually exserted at maturity, pubescence increasing in density
distally within each internode. Sessile spikelets (4.1)4.4-4.6(5) mm; callus
hairs 1-1.5 mm; keels of lower glumes scabrous only above the midpoint;
awns 2-11 mm; anthers 1, 1.4-2.4 mm, red. Pedicellate spikelets
vestigial or absent. 2n = 20.
Andropogon brachystachyus grows in sandy, often seasonally wet soils
of flatwoods, savannahs, pond margins, and scrublands of the southeastern United
States. It sometimes forms large populations, but does not invade disturbed
sites as do some morphologically similar forms of A.
virginicus var. virginicus.
12. Andropogon virginicus L.
Broomsedge Bluestem
Plants cespitose, dense and cylindrical to obpyramidal above. Culms
40-210 cm; internodes glaucous or not; branches erect to ascending,
usually straight, sometimes arching. Sheaths usually smooth, rarely somewhat
scabrous; ligules 0.2-1 mm, ciliate, cilia 0.2-1.3 mm; blades
11-52 cm long, 1.7-6.5 mm wide, smooth and glabrous or sparsely to densely pubescent
with spreading hairs. Inflorescence units 6-195 per culm; subtending
sheaths (2.1)3.1-4.6(6.7) cm long, (1.7)3-3.8(5.6) mm wide; peduncles
usually (1)4-6(30) mm, with 2-7 rames; rames (0.5)1.7-2.8(4.4) cm, sometimes
exserted at maturity, pubescence sparse basally and increasing in density distally
within each internode. Sessile spikelets (2.6)3.5-3.8(4.7) mm; callus
hairs 1-3 mm; keels of lower glume usually smooth below midlength,
scabrous distally; awns 6-21 mm; anthers 1(3), 0.6-1.5 mm, yellow
or purple. Pedicellate spikelets vestigial to absent. 2n = 20.
Andropogon virginicus is native from the southeastern United States to
northern South America, but has become established outside its native range in California, Hawaii, Japan, and Australia.
Three varieties are recognized, two of which contain morphologically distinct
variants. Andropogon virginicus hybridizes with A.
glomeratus and A. longiberbis
(Campbell 1986).
1 |
Leaves bluish-green, more or less strongly
glaucous ..... var. glaucus |
Leaves green, sometimes somewhat glaucous (2) |
|
Sheaths subtending the inflorescence units
(1.7)2.4-3.1(4) mm wide; inflorescences units usually with 2 rames; rames
(1.3)1.5-2.3(3) cm long; peduncles (1) 4-9 (30) mm long ..... var. decipiens
|
|
Sheaths subtending the inflorescences units (2.2)3.3-4.4(5.6)
mm wide; inflorescence units with 2-5(7) rames; rames (0.5)1.9-3.3(4.4)
cm long; peduncles (2)3-6(12) mm long ..... var. virginicus |
Andropogon virginicus var. decipiens C.S. Campb.
Culms 70-170 cm. Leaves green, sometimes slightly
glaucous. Inflorescence units usually with 2 rames; subtending sheaths
(1.7)2.4-3.1(4) mm wide; peduncles (1) 4-9 (30) mm; rames (1.3)1.5-2.3(3)
cm.
Andropogon virginicus var. decipiens grows in flatwoods, scrublands,
and disturbed sites, such as roadsides and cleared timberlands, of the southeastern
coastal plain.
Andropogon virginicus var. glaucus Hack.
Culms 60-180 cm. Leaves bluish-green, more or
less strongly glaucous. Inflorescence units with 2(3) rames; subtending
sheaths (2.7)3.1-3.8(5.5) mm wide; peduncles (2)3-4(10) mm; rames
(1.4)1.7-3(4) cm, not exserted.
Andropogon virginicus var. glaucus grows on moist or dry soils
of the coastal plain, from southern New Jersey to eastern Texas. Plants growing
on sandy, well-drained soils differ from those on poorly drained slopes in being
glabrous (rather than pubescent) beneath the subtending sheaths of the inflorescence
units, and in tending to have shorter rames.
Andropogon virginicus L. var. virginicus
Culms 40-210 cm; internodes green. Leaves green or slightly
glaucous, glabrous or pubescent, at least on the margins near the collar. Inflorescence
units with 2-5(7) rames; subtending sheaths (2.2)3.3-4.4(5.6) mm
wide; peduncles (2)3-6(12) mm; rames (0.5)1.9-3.3(4.4) cm, not
exserted.
Andropogon virginicus var. virginicus is the widespread and weedy
variety of A. virginicus that grows as a native species from the central
plains through Mexico and Central America to Colombia and, as a naturalized
species, in California, Hawaii, Japan, and Australia. Plants colonizing openings
in mature vegetation created by disturbance have green culms and green, pubescent
leaves. Those growing in poorly drained soils of pond margins, swales, and cutover
flatwoods have glaucous culms and glabrous, green to somewhat glaucous leaves.
Glaucous plants of A. virginicus var. virginicus differ from those
of var. decipiens in having no
exposed rames and, often, wider sheaths subtending the inflorescence units.
13. Andropogon longiberbis Hack.
Hairy Bluestem
Plants cespitose; cylindrical to oblong and more or less open in the
upper portion. Culms 50-100(150) cm; internodes green, sometimes
somewhat glaucous just below the node; branches mostly erect, straight.
Sheaths not scabrous; ligules 0.2-0.6 mm, ciliate, cilia 0.3-0.6
mm; blades 11-50 cm long, 2-5.5 mm wide, sparsely to densely pubescent,
most hairs appressed. Inflorescence units 7-97 (usually about 45) per
culm; subtending sheaths (2.5)3-4.5(6) cm long, (2.5)3.2-4.1(5.5) mm
wide; peduncles (1)3-4(13) mm, with 2(3) rames; rames (1.3)1.8-2.6(4)
cm, not exserted at maturity, pubescence increasing in density distally within
each internode. Sessile spikelets (3.5)4.1-4.5(5) mm; callus hairs
1.5-5 mm; keels of lower glumes scabrous only above the midpoint; awns
10-21 mm; anthers 1, 0.9-1.6 mm, yellow. Pedicellate spikelets
vestigial or absent. 2n = 20.
Andropogon longiberbis grows in sandy or rocky soils of roadsides, dunes,
sandhills, pinelands, and fields, from the southeastern United States to the
Bahamas. It resembles A. tracyi, but
usually differs in having more densely pubescent blades and a less slender appearance.
Andropogon longiberbis appears to hybridize with both
A. virginicus var. virginicus
and A. glomeratus var.
pumilus.
14. Andropogon glomeratus (Walter) Britton, Sterns &
Poggenb.
Bushy Bluestem, Bushy Beardgrass
Plants cespitose, upper portion dense, oblong to oblanceolate or obpyramidal.
Culms 20-250 cm; internodes green, sometimes glaucous; branches
mostly erect, straight. Sheaths usually scabrous, sometimes smooth; ligules
0.6-2.2 mm, sometimes ciliate, cilia to 0.9 mm; blades 13-109 cm long,
2.9-9.5 mm wide, glabrous or sparsely to densely pubescent, hairs usually spreading,
rarely appressed. Inflorescence units 10-600 per culm; subtending
sheaths (2.0)2.9-4.4(6.5) cm long, (1.5)2.3-3.4(4.4) mm wide; peduncles
(1)6-14(60) mm, with 2(4) rames; rames (1)1.7-2.5(3.5) cm, exserted or
not at maturity, pubescence sparse basally and increasing in density distally
within each internode. Sessile spikelets 3-5 mm; callus hairs
1-2.5 mm; keels of lower glumes sometimes scabrous below midlength, usually
scabrous distally; awns 6-19 mm; anthers 1(3), 0.5-1.5 mm, yellow,
red, or purple. Pedicellate spikelets vestigial or absent, sterile. 2n
= 20.
Andropogon glomeratus hybridizes with both A.
longiberbis and A. virginicus.
Some of its varieties are morphologically similar to the latter species.
1 |
Blades glaucous, glabrous, and smooth .....
var. glaucopsis |
Blades green, often pubescent or scabrous (2) |
|
Sheaths subtending the inflorescence units
1.5-3 mm wide; leaf sheaths usually smooth; ligules ciliate, the cilia 0.2-0.9
mm long ..... var. pumilus |
|
Sheaths subtending the inflorescence units (1.5)2.3-3.4(4.4)
mm wide; leaf sheaths often scabrous; ligules, when ciliate, with the cilia
no more than 0.5 mm long (3) |
|
Keels of the lower glumes
scabrous below and beyond midlength ..... var. scabriglumis |
|
Keels of the lower glumes usually smooth below midlength,
scabrous distally (4) |
|
Upper portion of the plants oblong to
obpyramidal; mature peduncles (4)11-35(60) mm long; anthers eventually falling
..... var. glomeratus |
|
Upper portion of the plants cylindrical to oblong; mature
peduncles 2-5(8) mm long; withered remnants of anthers retained within the
spikelets ..... var. hirsutior |
Andropogon glomeratus var. glaucopsis (Elliott)
C. Mohr
Plants usually oblong in the upper portion, sometimes cylindrical. Culms
1.2-2.2 m; internodes glaucous. Sheaths (1.3)2.3-3.1(4.7) mm wide,
usually smooth; ligules ciliate or not, cilia to 0.2 mm; blades
glaucous, glabrous, smooth. Subtending sheaths of inflorescence units usually
2-2.5 mm wide; peduncles shorter than 10 mm; rames usually shorter
than 2 cm, not exserted at maturity. Keels of lower glumes usually scabrous
beyond midlength.
Andropogon glomeratus var. glaucopsis grows in flatwoods, bogs,
ditches, swamps, pond margins, and swales of the southeastern coastal plain.
Andropogon glomeratus (Walter) Britton, Sterns & Poggenb.
var. glomeratus
Plants oblong to pyramidal in the upper portion. Culms 60-160
cm; internodes not glaucous. Sheaths (2)2.5-3.4(4.7) mm wide,
scabrous; ligules sometimes ciliate, cilia to 0.3 mm; blades green,
usually scabrous. Subtending sheaths of inflorescence units usually 2.5-3.4
mm wide; peduncles (4)11-35(60) mm; rames usually 2.1-2.9 cm,
exserted. Keels of lower glumes usually smooth below midlength, scabrous
distally; anthers eventually deciduous.
Andropogon glomeratus var. glomeratus grows in bogs, swamps, savannahs,
flatwoods, and ditches of the southeastern United States.
Andropogon glomeratus var. hirsutior (Hack.) C.
Mohr
Plants usually oblong in the upper portion, sometimes cylindrical. Culms
1-2 m; internodes not glaucous. Sheaths scabrous; ligules
ciliate or not, cilia to 0.3 mm; blades green, usually pubescent. Subtending
sheaths of inflorescence units (2)2.4-3.1(4) mm wide; peduncles 2-5(8)
mm; rames usually 1.7-2.8 cm, not exserted at maturity. Keels of lower
glumes usually smooth below midlength, scabrous distally; anthers
often retained within the spikelets.
Andropogon glomeratus var. hirsutior grows in ditches, swales,
bogs, flatwoods, and savannahs of the southeastern coastal plain, often forming
very large populations in cleared, low ground.
Andropogon glomeratus var. pumilus (Vasey) L.H.
Dewey
Plants oblanceolate to obpyramidal in the upper portion. Culms
to 2.5 m, but as short as 20 cm in poor soils; internodes not glaucous.
Leaf sheaths usually smooth; ligules ciliate, cilia 0.2-0.9 mm;
blades green, smooth or pubescent. Subtending sheaths of inflorescence
units (2)2.9-4.3(5.2) cm long, 1.5-3 mm wide; peduncles (2)8-15(40)
mm; rames 1.3-3 cm, exserted. Keels of lower glumes usually smooth
below midlength, scabrous distally; anthers often retained within the
spikelets.
Andropogon glomeratus var. pumilus is weedy and grows in disturbed,
wet or moist sites. It is abundant and widespread, extending from the southern
United States through Central America to northern South America.
Andropogon glomeratus var. scabriglumis C.S. Campb.
Plants oblanceolate to obpyramidal in the upper portion. Culms
80-150 cm; internodes not glaucous. Sheaths usually scabrous;
ligules ciliate, cilia 0.2-0.5 mm; blades green, smooth or pubescent.
Subtending sheaths of inflorescence units (2.3)2.9-4.5(6.3) cm long,
(1.5)2.3-3.3(4.4) mm wide; peduncles (2)5-10(16) mm; rames (1.7)1.9-2.3(2.8)
cm, exserted. Keels of lower glumes usually scabrous below and above
the midpoint.
Andropogon glomeratus var. scabriglumis grows in moist soils of
seepage slopes and the edges of springs, from California to New Mexico and southward
into Mexico.