ACHNATHERUM

Contributor: Mary E. Barkworth

ACHNATHERUM P. Beauv. , Essai Agrostogr. 19-20 (1812). Lasiagrostis Link, Hortus berol. 1:99 (1827). LEC: Achnatherum calamagrostis (L.) P. Beauv. [Niles & Chase, Contr. U.S. Natl. Herb. 246:160 (1925)].

Eriocoma Nutt., Gen. N. Amer. pl. 1:40 (1818). TYPE: Eriocoma cuspidata Nutt. [=Achnatherum hymenoides (Roemer & Schultes) Barkworth].

Fendleria Steudel, Syn. pl. glumac. 1:419 (1854). TYPE: Fendleria rhynchelytroides Steudel [=Achnatherum hymenoides (Roemer & Schultes) Barkworth].

Achnatherum aridum (M.E. Jones) Barkworth, Phytologia 74:6 (1993). Stipa arida M.E. Jones, Proc. Calif Acad. Sci., Ser. 2, 5:725 (1895). HOL: POM 116537!;U.S.A., Utah: Marysvale, on very dry talus slopes in shingle; 4 Jun 1894; M.E. Jones 5377.

Stipa mormonum Mez , Repert. Spec. Nov. Regni Veg. 17:209 (1921). TYPE: U.S.A., Utah; M.E. Jones 2106.

Achnatherum ×bloomeri (Bol.) Barkworth (Bol.) Barkworth, Phytologia 74:14 (1993). Stipa bloomeri Bol. , Proc. Calif. Acad. Sci. 4:168 (1872); Oryzopsis bloomeri (Bol.) Ricker in Piper, Contr. U.S. Natl. Herb. 11:109 (1906); ×Stiporyzopsis bloomeri (Bol.) B.L. Johnson, Amer. J. Bot. 32:602 (1945). HOL: CAS 1145; U.S.A., California: Bloody Canyon, near Mono Lake; H.N. Bolander 6116. [Achnatherum hymendoides × Achnatherum occidentale, fide B.L. Johnson, Amer. J. Bot. 32:599-608].

Achnatherum caudatum (Trin.) S.W.L. Jacobs & J. Everett, Telopea 6:582 (1996). Stipa caudata Trin., Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 6, Sci. Math. 1:75 (1830). HOL: LE; Chile; J. Lindley.

Stipa brachychaeta Godron, Mém. Sect. Sci. Acad. Sci. Montpellier 1:450 (1853); Achnatherum brachychaetum (Godron) Barkworth, Phytologia 74:6 (1993). TYPE: France: foreign plants growing in Port Juvénale, near Montpellier.

Stipa bertrandii [as "bertrandi"] Phil., Linnaea 33:283 (1864). TYPE: Chile: at the foot of the Andes, near Santiago; C.E.Bertrand.

Stipa amphicarpa Phil. Anales Univ. Chile 93:726 (1896) [fide Matthei, Gayana, Bot. 13:69 (1965). TYPE: Chile: Angol; Dec 1887; R.A. Philippi.

Stipa litoralis Phil. Anales Univ. Chile 93:717 (1896). TYPE: Chile: Quiriquina Island; R.A. Philippi.

Stipa clandestina Hackel, Repert. Spec. Nov. Regni Veg. 8:516 (1910); Achnatherum clandestinum (Hackel) Barkworth, Phytologia 74:6 (1993). LEC: US 1030718; México: near Michoacan; G. Arsè ne 3441 [Hitchc., Contr. U.S. Natl. Herb. 247:238 (1925)].

Achnatherum contractum (B.L. Johnson) Barkworth, Phytologia 74:6 (1993). Oryzopsis hymenoides var. contracta B.L. Johnson, Bot. Gaz. 107:24 (1945); Oryzopsis contracta Shechter in Shechter & B.L. Johnson, Brittonia 18:342 (1966). HOL: RM!; U.S.A., Wyoming: Carbon County, Freezeout Hills; E. Nelson 4850.

Achnatherum coronatum (Thurber) Barkworth, Phytologia 74:6 (1993). Stipa coronata Thurber in S. Watson, Bot. California 2:287 (1880). SYN: [1] GH 17897!; U.S.A., California: southern California; 1876; C.C. Parry and J.G. Lemmon 422; [2] GH 17898; U.S.A., California: moist hillsides, Julian City, San Diego; H.N. Bolander.

Achnatherum curvifolium (Swallen) Barkworth, Phytologia 74:7 (1993). Stipa curvifolia Swallen, J. Wash. Acad. Sci. 23:456 (1933). HOL: US 1538063!; U.S.A., New Mexico: Eddy County, Guadalupe Mountains, near mouth of North Fork of Rocky Arroyo, in crevices of limestone cliff; 29 Apr 1932; H. Wilkins 1660.

Achnatherum depauperatum (M.E. Jones) Barkworth, ined. Stipa parishii var. depauperata M.E. Jones, Contr. W. Bot. 14:11 (1912); Stipa coronata var. depauperata (M.E. Jones) Hitchc., J. Wash. Acad. Sci. 24:292 (1934); Achnatherum parishii subsp. depauperatum (M.E. Jones) Barkworthdepauperatum (M.E. Jones) Barkworth, Phytologia 74:11 (1993).. TYPE: U.S.A., Utah: Detroit [a now-vanished mining town]; 28 May 1891; M.E. Jones.

Achnatherum diegoense (Swallen) Barkworth, Phytologia 74:7 (1993). Stipa diegoensis Swallen, J. Wash. Acad. Sci. 30:212-213 (1940). HOL: US 1761177!; U.S.A., California: San Diego County, Proctor Valley, near Jamul, along vernal stream in chaparral; 23 May 1938; F.F. Gander 5778.

Achnatherum eminens (Cav.) Barkworth, Phytologia 74:7 (1993). Stipa eminens Cav., Icon. 5:42, pl. 467, f. 1 (1799). TYPE: México: Chalma.
Stipa erecta E. Fourn., Mexic. pl. 2:75 (1886), non Trin. (1824). TYPE: México: Tehuacan; F.M. Liebmann 654.

Stipa flexuosa Vasey , Bull. Torrey Bot. Club 15:49 (1888). TYPE: U.S.A., Texas: Presidio County, Chenate Mountains; 1887; G.C. Nealley 710.

Achnatherum hendersonii (Vasey) Barkworth, Phytologia 74:7 (1993). Oryzopsis hendersonii Vasey, Contr. U.S. Natl. Herb. 18:267 (1893); Oryzopsis exigua var. hendersonii (Vasey) M.E. Jones, Contr. W. Bot. 14:11 (1912); Stipa hendersonii (Vasey) Mehlenb. in Spellenb. & Mehlenb., Canad. J. Bot. 49:1568 (1971). TYPE: U.S.A., Washington: Clements Mt., near North Yakima; L.F. Henderson 2249.

Achnatherum hymenoides (Roemer & Schultes) Barkworth, Phytologia 74:7 (1993). Stipa hymenoides Roemer & Schultes, Syst. veg. 2:339 (1817); Oryzopsis hymenoides (Roemer & Schultes) Ricker in Piper, Contr. U.S. Natl. Herb. 11:109 (1906); Eriocoma hymenoides (Roemer & Schultes) Rydb., Bull. Torrey Bot. Club 39:102 (1912). Stipa membranacea Pursher & Schultes) Rydb., Bull. Torrey Bot. Club 39:102 (1912). Stipa membranacea Pursh , Fl. Amer. sept. 2:728 (1813) [title page 1814], nom. illeg., non L. (1753); Oryzopsis membranacea Vasey, U.S.D.A. Div. Bot. Bull. 122:pl. 10 (1891), nom. superfl.; Eriocoma membranacea Beal, Grasses N. Amer. 2:232 (1896), nom. illeg. et superfl., non Steudel (1840). HOL: PH!; U.S.A.: banks of the Missouri River; E.P. Bradbury.

Eriocoma cuspidata Nutt., Gen. N. Amer. pl. 1:40 (1818); Milium cuspidata [as "cuspidatum"] (Nutt.) Sprengel, Syst. veg. 1:251 (1824) [title page 1825]; Oryzopsis cuspidata (Nutt.) Bentham ex Vasey, Dept. Agric. Special Rep. 63:23 (1883). HOL: PH!; U.S.A.: Platte plains; T. Nuttall.

Urachne lanata Trin., Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math. 3:126 (1834). Eriocoma membranacea Steudel, Nomencl. bot., ed. 2, 1:586 (1840). TYPE: North America.

Fendleria rhynchelytroides Steudel, Syn. pl. glumac. 1:420 (1854). HOL: GH 23719!; U.S.A., New Mexico[: probably Santa Fe County, near Santa Fe]; 1847; A. Fendler 979.

Achnatherum latiglume (Swallen) Barkworth, Phytologia 74:8 (1993). Stipa latiglumis Swallen, J. Wash. Acad. Sci. 23:198, f. 1 (1933). HOL: US 992334!; U.S.A., California: Yosemite Valley, Camp Lost Arrow; 22 Jun 1911; L. Abrams 4469.

Achnatherum lemmonii (Vasey) Barkworth, Phytologia 74:8 (1993). HOL: US 556900!; U.S.A., California: Plumas County, Mohawk Valley; May 1889; J.G. Lemmon 5456.

Achnatherum lemmonii (Vasey) Barkworth subsp. lemmonii, Phytologia 74:8 (1993). Stipa pringlei var. lemmonii Vasey, Contr. U.S. Natl. Herb. 31:55 (1892); Stipa lemmonii (Vasey) Scribner, Circ. Div. Agrostol. U.S.D.A. 30:3 (1901). HOL: US 556900!; U.S.A., California: Plumas County, Mohawk Valley; May 1889; J.G. Lemmon 5456.

Stipa columbiana Macoun, Cat. Canad. pl. 24:191 (1888), nom. rejic.[Barkworth & Maze, Taxon 28:621-624 (1979)], non auct. pl. LEC: CAN 9899!; Canada, British Columbia: Yale; 17 May 1875; J.M. Macoun, Geol. Survey Can. 28940 [Hitchc., Contr. U.S. Natl. Herb. 247:253 (1925)].
Stipa lemmonii var. jonesii Scribner, Circ. Div. Agrostol. U.S.D.A. 30:4 (1901). HOL: US 556899!; U.S.A., California: Emigrant Gap; 28 Jun 1882; M.E. Jones 3298.

Achnatherum lemmonii subsp. pubescens (Crampton) Barkworth , Phytologia 74:8 (1993). Stipa lemmonii var. pubescens Crampton, Leafl. W. Bot. 7:220 (1955). HOL: DAV 21077!; U.S.A., California: Tehama County, Whitlock Camp-Round Mountain area west of Paskenta, serpentine slopes in chaparral, elev. about 4000 feet; 16 Jun 1954; B.S. Crampton 2000.

Achnatherum lettermanii (Vasey) Barkworth, Phytologia 74:9 (1993). Stipa lettermanii Vasey, Bull. Torrey Bot. Club 13:53 (1886); Stipa viridula var. lettermanii (Vasey) Vasey, Contr. U.S. Natl. Herb. 31:50 (1892). HOL: US 556904!; U.S.A., Idaho: Snake River; Aug 1885; G.W. Letterman 102.
Stipa viridula var. minor Vasey, Contr. U.S. Natl. Herb. 31:50 (1892); Stipa minor (Vasey) Scribner, Bull. Div. Agrostol. U.S.D.A. 112:46 (1898). LEC: US 556903!; U.S.A., Colorado: Kelso Mountain, near Torrey's Peak, alt. 13,000 feet; 13 Aug 1889; G.W. Letterman 95 [Hitchc., Contr. U.S. Natl. Herb. 247:253 (1925)].

Achnatherum lobatum (Swallen) Barkworth, Phytologia 74:9 (1993). Stipa lobata Swallen, J. Wash. Acad. Sci. 23:199, f. 2 (1933). HOL: US 905722!; U.S.A., New Mexico: Guadalupe Mountains, Queen Ranger Station, on rocky hill, alt. 6000-7000 feet; 3-6 Sep 1915; A.S. Hitchcock 13502.

Achnatherum nelsonii (Scribner) Barkworth, Phytologia 74:9 (1993). HOL: US 556901!; U.S.A., Wyoming: Albany County, Wood's Landing; 9 Aug 1897; A. Nelson 3963.

Achnatherum nelsonii (Scribner) Barkworth subsp. nelsonii, Phytologia 74:9 (1993). Stipa nelsonii Scribner , Bull. Div. Agrostol. U.S.D.A. 112:46 (1898); Stipa columbiana var. nelsonii (Scribner) Hitchc., Contr. U.S. Natl. Herb. 247:254 (1925); Stipa occidentalis var. nelsonii (Scribner) C.L. Hitchc. in C.L. Hitchc., Cronquist & Ownbey, Vascular Plants of the Pacific Northwest 1:715 (1969).HOL: US 556901!; U.S.A., Wyoming: Albany County, Wood's Landing; 9 Aug 1897; A. Nelson 3963.
Stipa williamsii Scribner, Bull. Div. Agrostol. U.S.D.A. 112:45 (1898). HOL: US 556907!; U.S.A., Wyoming: west side of Big Horn Mountains, near Monument Spring, dry soil; 3 Aug 1897; T.A. Williams 2804.

Achnatherum nelsonii subsp. dorei (Barkworth & Maze) Barkworth (Barkworth & Maze) Barkworth, Phytologia 74:9 (1993). Stipa nelsonii subsp. dorei Barkworth & Maze Barkworth & Maze, Taxon 28:623 (1979); Stipa nelsonii var. dorei (Barkworth & Maze) Dorn, Vascular Plants of Wyoming 298 (1988); Achnatherum nelsonii var. dorei(Barkworth & Maze) Dorn, Vascular Plants of Wyoming, ed. 3, 376 (2001). HOL: DAO 168288!; Canada, Alberta: Dungarvan Creek; W.G. Dore 12136.

Stipa columbiana auct. pl. auct. pl., non Macoun (1888).

Stipa occidentalis var. minor sensu C.L. Hitchcock in C.L. Hitchc., Cronquist & Ownbey, Vascular Plants of the Pacific Northwest 1:714 (1969).

Achnatherum nevadense (B.L. Johnson) Barkworth, Phytologia 74:9 (1993). Stipa nevadensis B.L. Johnson, Amer. J. Bot. 49:257 (1962). HOL: UC!; U.S.A., California: Mono County, Upper Twin Lake, near Bridgeport; B.L. Johnson 211.

Achnatherum occidentale (Thurber) Barkworth, Phytologia 74:10 (1993). HOL: GH 22338!; U.S.A., California: Yosemite Trail; 20 Aug 1866; H.N. Bolander 5038.

Achnatherum occidentale (Thurber) Barkworth subsp. occidentale, Phytologia 74:10 (1993). Stipa occidentalis Thurber in S. Watson, Botany (fortieth parallel) 5:380 (1871). Stipa occidentalis var. montana Merr. & Burtt Davymontana Merr. & Burtt Davy in H.M. Hall, Univ. Calif. Pub. Bot. 1:62 (1902). HOL: GH 22338!; U.S.A., California: Yosemite Trail; 20 Aug 1866; H.N. Bolander 5038.

Achnatherum occidentale subsp. californicum (Merr. & Burtt Davy) Barkworth, Phytologia 74:10 (1993). Stipa californica Merr. & Burtt Davy in H.M. Hall, Univ. Calif. Pub. Bot. 1:61 (1902); Stipa occidentalis var. californica (Merr. & Burtt Davy) C.L. Hitchc.californica (Merr. & Burtt Davy) C.L. Hitchc. in C.L. Hitchc., Cronquist &Ownbey, Vascular Plants of the Pacific Northwest 1:715 (1969). TYPE: U.S.A., California: San Jacinto Mountains, north side of Fuller's Ridge; Jul 1901; H.M. Hall 2556.

Stipa nelsonii var. longiaristata Barkworth & Maze, Taxon 28:623 (1979); Achnatherum nelsonii subsp. longiaristatum (Barkworth & Maze) Barkworthlongiaristatum (Barkworth & Maze) Barkworth, Phytologia 74:9 (1993). HOL: WS!; U.S.A., Washington: 8-9 miles west of Spokane; J.R. Swallen 6231.

Achnatherum occidentale subsp. pubescens (Vasey) Barkworth , Phytologia 74:10 (1993). Stipa viridula var. pubescens Vasey, Contr. U.S. Natl. Herb. 31:50 (1892); Stipa occidentalis var. pubescens (Vasey) Roy L. Taylor & MacBryde, Canad. J. Bot. 56:193 (1978). Stipa elmeri Piper & Brodie ex Scribner, Bull. Div. Agrostol. U.S.D.A. 112:46 (1898). TYPE: U.S.A., Washington: Columbia River; 1883; W.N. Suksdorf.

Achnatherum parishii (Vasey) Barkworth, Phytologia 74:11 (1993). Stipa parishii Vasey , Bot. Gaz. 7:33 (1882); Stipa coronata subsp. parishi (Vasey) Hitchc., Contr. U.S. Natl. Herb. 247:227 (1925). HOL: US 74509!; U.S.A., California: San Bernardino Mountains; S.B. Parish 1079.

Achnatherum perplexum Hoge & Barkworth, Phytologia 74:11 (1993). Stipa perplexa (Hoge & Barkworth) Wipff & S.D. Jones, Phytologia 77:461 (1994). HOL: US!; U.S.A., New Mexico: Bernallilo County, 9 miles south of Tijeres on NM Route 14; 8 Sep 1985; M.E. Barkworth 4764.

Achnatherum pinetorum (M.E. Jones) Barkworth, Phytologia 74:12 (1993). Stipa pinetorum M.E. Jones, Proc. Calif. Acad. Sci., Ser. 2, 5:724 (1895). NEO: US 236788!; U.S.A., Utah: Panguitch Lake; 8 Sep 1894; M.E. Jones 6023p [Barkworth, Phytologia 74:12 (1993)].

Achnatherum richardsonii (Link) Barkworth, Phytologia 74:12 (1993). Stipa richardsonii Link , Hortus berol. 2:245 (1833); Oryzopsis richardsonii (Link) Beal, Bot. Gaz. 15:111 (1890). HOL: B [lost, presumably destroyed]; cultivated at botanical garden in Berlin, Germany from seed sent by [J.] Richardson.

Achnatherum robustum (Vasey) Barkworth, Phytologia 74:12 (1993). Stipa viridula var. robusta Vasey in J.M. Coulter, Contr. U.S. Natl. Herb. 12:56 (1890); Stipa robusta (Vasey) Scribner in Rydb. & Shear, Bull. Div. Agrostol. U.S.D.A. 5:23 (1897), nom. cons. Stipa vaseyi Scribner , Bull. Div. Agrostol. U.S.D.A. 112:46 (1898), nom. superfl. HOL: US 993051!; U.S.A., New Mexico; G.R. Vasey, typ. cons. [Barkworth, Taxon 42:711-713 (1993)].

Achnatherum scribneri (Vasey) Barkworth, Phytologia 74:13 (1993). Stipa scribneri Vasey , Bull. Torrey Bot. Club 11:125 (1884). HOL: US 556905!; U.S.A., New Mexico: Santa Fe, dry hillsides; G.R. Vasey.

Achnatherum splendens (Trin.) Nevski, Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyss. Rast. 4:224 (1937). Stipa splendens Trin. in Sprengel, Neue Entd. 2:54 (1821); Lasiagrostis splendens (Trin.) Kunth, Révis. gramin. 1:58 (1829). HOL: LE (Fischer Herb., sub "Agrostis longiaristata"); Russia: "Transbaikal" [Siberia].

Achnatherum stillmanii (Bol.) Barkworth, Phytologia 74:14 (1993). Stipa stillmanii Bol. , Proc. Calif. Acad. Sci. 4:169 (1872). HOL: GH 17890!; U.S.A., California: Sierra Nevada, Blue Canyon; Jul 1870; H.N. Bolander.

Achnatherum swallenii (C.L. Hitchc. & Spellenb.) Barkworth, Phytologia 74:14 (1993). Oryzopsis swallenii C.L. Hitchc. & Spellenb., Brittonia 20:164 (1968). HOL: WTU 227273!; U.S.A., Idaho: Clark County, open sagebrush slope just north of Birch Creek; 7 Jul 1965; C.L. Hitchcock 23868.

Achnatherum thurberianum (Piper) Barkworth, Phytologia 74:14 (1993). Stipa thurberiana Piper , Circ. Div. Agrostol. U.S.D.A. 275:10 (1900). Stipa occidentalis Thurber in Torrey in Wilkes, U.S. Expl. Exped. 17:483 (1874). HOL: US 556917; U.S.A., Washington: north branch of the Columbia [River]; C. Pickering and W.D. "Breckenridge" [Brackenridge].

Achnatherum wallowaense Maze & K.A. Robson, Madroño 43:401 (1996). HOL: US; U.S.A., Oregon: Wallowa County, Wallowa-Whitman National Forest, ca. 34 km north of Enterprise, near Boner Gulch along Forest Service Road 46, 1481 m, 45° 43' 41.16" N 117° 8' 10.32" W; 26 Jun 1993; J. Maze 1007, E. Maze, K.A. Robson, and T. Henn.

Achnatherum webberi (Thurber) Barkworth, Phytologia 74:13 (1993). Eriocoma webberi Thurber in S. Watson, Bot. California 2:283 (1880); Oryzopsis webberi (Thurber) Bentham ex Vasey, Dept. Agric. Special Rep. 63:23 (1883); Stipa webberi (Thurber) B.L. Johnson, Bot. Gaz. 107:25 (1945). TYPE: U.S.A., California: Sierra Valley; May 1871; H.N. Bolander.