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This page has been developed in partial response to requests at a workshop held in conjunction with the  Wheat Genetics Symposium in Saskatoon, 1998.  It gives the synonymy for the species of Aegilops that have been found in North America north of Mexico, based on van Slageren (1994).  Eventually, I hope to link the listed species to images that will help in identification.  For now, admire the odd mixture that is on the right. 

A synopsis of the genus and the genomic constitution of its species based on Dvorak (1998) is given on a separate page.  That page will also, one day, include a synopsis of the native distribution of the individual species.  For really great distribution maps, see van Slageneren (1994).   They illustrate very clearly the concept the  Fertile Crescent fro m the point of view of plants. 

Names in bold are accepted; names in italics are synonyms.  This material is not copyrighted (in other words, you are welcome to copy it).Citations of authorities and literature follows international standards.  The text was largely prepared and edited by Dr. Kathleen Capels in connection with her work as Associate Editor of the Manual.  
Mary Barkworth

AEGILOPS L., Sp. pl. 2:1050 (1753), nom. cons. prop. Aegilopodes A. Löve, Biol. Zentralbl. 101:207 (1982). LEC: Aegilops triuncialis L., typ. cons. prop. [Bowden, Canad J. Bot. 37:675 (1959)]. [Also see M.W. van Slageren, Wild Wheats: A Monograph of Aegilops L. and Amblyopyrum (Jaub. & Spach) Eig (1994)-the source for part of this synonymy-for a more complete treatment of synonymy of Aegilops.]

Aegilops subg. Cylindropyrum Jaub. & Spach, Ill. pl. orient. 4:12 (1850); Cylindropyrum (Jaub. & Spach) A. Löve, Biol. Zentralbl. 101:206 (1982). TYPE: Cylindropyrum cylindricum (Host) A. Löve [=Aegilops cylindrica Host]

Aegilops subg. Gastropyrum Jaub. & Spach, Ill. pl. orient. 4:17 (1850); Gastropyrum (Jaub. & Spach) A. Löve, Biol. Zentralbl. 101:208 (1982). TYPE: Gastropyrum ventricosum (Tausch) A. Löve [=Aegilops ventricosa Tausch]

Patropyrum A. Löve, Biol. Zentralbl. 101:206 (1982). TYPE: Patropyrum tauschii (Cosson) A. Löve [=Aegilops tauschii Cosson]

Aegilops crassa Boiss., Diagn. pl. orient., Ser. 1, pt. 7:129 (1846). Triticum crassum (Boiss.) Aitch. & Hemsley in Aitch., Trans. Linn. Soc. London, Bot., Ser. 2, 3:127 (1886); Gastropyrum crassum (Boiss.) A. Löve, Feddes Repert. 95:501 (1984). HOL: G (G-BOIS); "Persia" [Iran]: Persepolis [Takht-e Jamshid]; 16 Apr 1842; K.G.T. Kotschy 248

Aegilops platyathera Jaub. & Spach, Ill. pl. orient. 4:17, pl. 313 (1851). Aegilops crassa ß macrathera Boiss., Fl. orient. 5:677 (1884); Triticum crassum var. macratherum (Boiss.) Thell., Fl. adv. Montpellier 150 (1912) HOL: P; "Mesopotamia" [Turkey/Syria/Iraq]: between Mardin and Mosul [Al-Mawsil]; P.M.R. Aucher-Éloy 2913.

Aegilops crassa var. flavescens Popova, Trudy Prikl. Bot. Selekts. 13:477 (1923). TYPE: "Turkestan" [Uzbekistan]: Syr-Darja [Syrdarja]; G.M. Popova.

 

Aegilops crassa var. fuliginosa Popova, Trudy Prikl. Bot. Selekts. 13:477 (1923); Aegilops crassa forma fuliginosa (Popova) K. Hammer, Feddes Repert 91:234 (1980). TYPE: "Turkestan" [Uzbekistan]: Syr-Darja [Syrdarja]; G.M. Popova.

Aegilops crassa var. rubiginosa Popova, Trudy Prikl. Bot. Selekts. 13:477 (1923); Aegilops crassa forma rubiginosa (Popova) K. Hammer, Feddes Repert 91:234 (1980). TYPE: "Turkestan" [Uzbekistan]: Syr-Darja [Syrdarja]; G.M. Popova.

Aegilops crassa var. glumiaristata Eig, Bull. Soc. Bot. Genčve, Ser. 2, 19:328 (1928). HOL: LE [not found]; "Turkestan" [Turkmenistan]: Amu-Daria [Amu-Darja]; 30 Apr 1915; M.G. Popov.

Aegilops cylindrica Host, Icones gram. austriac. 2:6, pl. 7 (1802). Triticum cylindricum (Host) Ces., Pass. & Gibelli, Comp. fl. ital. 14:86 (1869); Triticum caudatum B. cylindricum (Host) Ach. & Graebner, Syn. mitteleur. Fl. 21:710 (1902); Aegilops caudata subsp. cylindrica (Host) Hegi, Ill. Fl. Mitt.-Eur. 1:390 (1908); Aegilops caudata ß [var.] cylindrica (Host) Fiori, Nuova Fl. Italia 1:160 (1923); Cylindropyrum cylindricum (Host) A. Löve, Biol. Zentralbl. 101:207 (1982). LEC: BP; [Hungary]: Budapest; P. Kitaibel 226 [Slageren, Wild Wheats 200 (1994)].

Aegilops cylindrica var. rumelica [as "rumelicum"] Velen., Fl. bulg. 627 (1891); Triticum cylindricum var. rumelicum (Velen.) Stoj. & Stef., Fl. Bulg. 1:169 (1924); Triticum cylindricum ß rumelicum (Velen.) Hayek in Hayek & Markgr., Repert. Spec. Nov. Regni Veg. Beih. 30, 3:228 (1932). HOL: PRC; [Bulgaria]: Philippopolis [Plovdiv]; Jun 1890; K. Keck and T. Pilcher.

Aegilops caudata var. hirsuta (Binz) Hegi, Ill. Fl. Mitt.-Eur. 1:390 (1908); Aegilops cylindrica var. hirsuta (Binz) Hegi & Suess., Ill. Fl. Mitt.-Eur., ed. 2, 1:500 (1936). TYPE: not designated.

Aegilops cylindrica pubescens Kloos in Jansen, Ned. Kruidk. Arch. 1914: 66 (1914); Aegilops cylindrica var. pubescens (Kloos) Jansen, Fl. Neerlandica 12:122 (1951). LEC: L 953.200-231; [Netherlands:] Wormerveer; 27 Jun 1914; A.W. Kloos [Slageren, Wild Wheats 202 (1994)].

Aegilops cylindrica var. albescens Popova, Trudy Prikl. Bot. Selekts. 13:476 (1923). TYPE: "Turkestan" [Uzbekistan]: Syr-Darja [Syrdarja], Samarkand; G.M. Popova.

Aegilops cylindrica var. brunnea Popova, Trudy Prikl. Bot. Selekts. 13:476 (1923); Aegilops cylindrica forma brunnea (Popova) K. Hammer, Feddes Repert 91:233 (1980). TYPE: "Turkestan" [Uzbekistan]: Syr-Darja [Syrdarja]; G.M. Popova.

Aegilops cylindrica var. ferruginea Popova, Trudy Prikl. Bot. Selekts. 13:476 (1923); Aegilops cylindrica forma ferruginea (Popova) K. Hammer, Feddes Repert 91:232 (1980). SYN: [1] [Russia:] Saratov; Vilenski; [2] "Turkestan" [Uzbekistan]: Syr-Darja [Syrdarja]; G.M. Popova.

Aegilops cylindrica var. flavescens Popova, Trudy Prikl. Bot. Selekts. 13:476 (1923); Aegilops cylindrica forma flavescens (Popova) K. Hammer, Feddes Repert 91:233 (1980). TYPE: "Turkestan" [Uzbekistan]: Syr-Darja [Syrdarja], Samarkand; G.M. Popova.

Aegilops cylindrica var. fuliginosa Popova, Trudy Prikl. Bot. Selekts. 13:476 (1923); Aegilops cylindrica forma fuliginosa (Popova) K. Hammer, Feddes Repert 91:233 (1980). TYPE: "Turkestan" [Uzbekistan]: Syr-Darja [Syrdarja]; G.M. Popova.

Aegilops cylindrica var. rubiginosa Popova, Trudy Prikl. Bot. Selekts. 13:477 (1923); Aegilops cylindrica forma rubiginosa (Popova) K. Hammer, Feddes Repert 91:233 (1980). TYPE: "Turkestan" [Uzbekistan]: Syr-Darja [Syrdarja], Samarkand; G.M. Popova.

Aegilops cylindrica subsp. aristulata Zhuk., Trudy Prikl. Bot. 18:507 (1928); Aegilops cylindrica var. aristulata (Zhuk.) Tzvelev, Novosti Sist. Vyssh. Rast. 10:37 (1973). SYN: [1] "Transcaucasia" [Azerbaijan]: Nakhichevan province; Kuleschov ; [2] [Russia:] Daghestan, near Derbent; Zherdeva; [3] Afghanistan: Chadjatchit-Charzar; Bukinitch.

Aegilops cylindrica var. pauciaristata Eig, Repert. Spec. Nov. Regni Veg. Beih. 55:103 (1929); Aegilops cylindrica subsp. pauciaristata (Eig) Chennav., Acta Hort. Gothob. 23:166 (1960); Cylindropyrum cylindricum subsp. pauciaristatum (Eig) A. Löve, Feddes Repert. 95:500 (1984). SYN: [1] G (G-BOIS), L, P; [Turkey:] Cappodicia, Caesareebene [Caesarea]; 1886; B. Balansa 42; [2] Romania: Focsani; 1923; O. Savulascu; [3] Serbia: east Serbia, near Prevlua; 1890; L. Adamovic; [4] BM, G, K, MO, PH, W; Bulgaria: near Sliven, Mt. Baramuk; 1907; C.K. Schneider 1426; [5] NY; "Russia" [Ukraine]: Cherson, near Limane; A. Rehmann 174.

Aegilops cylindrica var. multiaristata Jansen & Wachter, Ned. Kruidk. Arch. 1932: 135, f. 5c, 138 (1932). HOL: L; Netherlands: Gorinchem; Jul 1912; J.T. Henrard.

Aegilops cylindrica var. prokhanovii Tzvelev, Novosti Sist. Vyssh. Rast. 10:37 (1973). HOL: LE; [Russia:] Daghestan, Machacz-Kala district, Mt. Narat-Tjube, Alti-Bujukski Pass; 22 Jun 1955; Y.I. Prokhanov 280.

Aegilops cylindrica var. kastorianum Karataglis, Pl. Syst. Evol. 163:19 (1989). HOL: TAU; [Greece:] Kastoria; S.S. Karataglis 9.

Aegilops geniculata Roth, Bot. Abh. Beobacht. 45 (1787). HOL: B (B-W): Germany; A.W. Roth. [Acceptance of this name is based on the assumption that the name Aegilops ovata L. will be rejected, either as a nom. ambig. [Greuter & Rech. f., Boissiera 13:1-206 (1967); Kerguélen, Bull. Soc. Bot. France 123:317-24 (1976)] or as a name that has persistently and consistently been used in a way that excludes its type [Lambinon, Taxon 30:361-66 (1981)]. At this time, the Committee has made no decision. Usage in this Manual conforms to that adopted in Flora Europaea, Flora of Turkey and the East Aegean Islands, and Flora Palćstina.]

Phleum aegylops Scop., Fl. carniol., ed. 2, 1:55 (1771). TYPE: [Italy:] near Trieste; J.A. Scopoli

Aegilops echinus Godron, Fl. juvenalis 48 (1853); Triticum ovatum var. echinus (Godron) Thell., Fl. adv. Montpellier 145 (1912); Aegilops ovata var. echinus (Godron) Eig, Repert. Spec. Nov. Regni Veg. Beih. 55:145 (1929); Aegilops geniculata var. echinus (Godron) K. Hammer, Feddes Repert. 91:241 (1980). HOL: NCY; France: foreign plants growing in Port Juvénale, near Montpellier; A. Touchy 294.

Aegilops brachyathera Pomel, Bull. Soc. Sci. Phys. Algérie 13:389 (1876) [dated 1875]; Triticum ovatum var. brachyathera ("Hackel, in litt. " [error for Pomel]) T. Durand & Schinz, Consp. fl. afric. 5:938 (1894); Aegilops ovata var. brachyathera (Pomel) Eig, Repert. Spec. Nov. Regni Veg. Beih. 55:145 (1929); Aegilops ovata subsp. brachyathera (Pomel) Chennav., Acta Horti Gothob. 23:164 (1960); Aegilops geniculata var. brachyathera (Pomel) K. Hammer, Feddes Repert. 91:241 (1980). Aegilops ovata var. eigiana Maire & Weiller in Maire, Flore de l'Afrique du Nord 3:369 (1955). HOL: AL; Algeria: Sersou, Aďn Toucria, Bourbaki; 14 May 1860; A.N. Pomel.

Aegilops fonsii Sennen, Bull. Soc. Bot. France 69:91 (1922). HOL: BC; Spain: Barcelona, Rubí; 5 Jun 1921; Frčre Sennen 4220.

Aegilops ovata subsp. gibberosa Zhuk., Trudy Prikl. Bot. 18:471 (1928); Aegilops geniculata subsp. gibberosa (Zhuk.) K. Hammer, Feddes Repert. 91:241 (1980). SYN: [1] France: southern France, Toulon; N.I. Vavilov; [2] Italy: Sardinia; N.I. Vavilov.

Aegilops ovata subsp. globulosa Zhuk., Trudy Prikl. Bot. 18:473 (1928); Aegilops geniculata subsp. globulosa (Zhuk.) A. Löve, Feddes Repert. 95:503 (1984). HOL: WIR 2723; Tunisia: near Maktar; 10 Aug 1926; N.I. Vavilov.

Aegilops ovata var. lanuginosa Zhuk., Trudy Prikl. Bot. 18:474 (1928); Aegilops geniculata var. lanuginosa (Zhuk.) K. Hammer, Feddes Repert. 91:241 (1980). SYN: [1] Spain; N.I. Vavilov; [2] Italy: Sicily; N.I. Vavilov.

Aegilops ovata subsp. planiuscula Zhuk., Trudy Prikl. Bot. 18:473 (1928). LEC: WIR 938; Spain: Morede, near Granada, near barley fields; 23 Jun 1927; N.I. Vavilov 55130 [Slageren, Wild Wheats 225 (1994)].

Aegilops ovata var. puberulla Zhuk., Trudy Prikl. Bot. 18:473 (1928); Aegilops geniculata var. puberulla (Zhuk.) K. Hammer, Feddes Repert. 91:240 (1980). SYN: Algeria; N.I. Vavilov; [2] Italy: Sicily; N.I. Vavilov.

Aegilops ovata subsp. umbonata Zhuk., Trudy Prikl. Bot. 18:472 (1928). LEC: WIR 960; "Palestine" [Israel]: upper Galilea; 6 Sep 1926; N.I. Vavilov 29512 [Slageren, Wild Wheats 225 (1994)].

Aegilops ovata var. vernicosa Zhuk., Trudy Prikl. Bot. 18:473 (1928). SYN: [1] Algeria: Algiers, Frenda; N.I. Vavilov 239; [2] Tunisia; N.I. Vavilov.

Aegilops ovata var. africana Eig, Repert. Spec. Nov. Regni Veg. Beih. 55:144 (1929); Aegilops geniculata var. africana (Eig) H. Scholz, Willdenowia 7:420 (1974). SYN: [1] Morocco: Casa Blanca [Casablanca]; 1871; Hackel; [2] Libya: Cyrneaica, Derra; 5 Apr 1887; P.H.W. Taubert 181.

Aegilops ovata subsp. atlantica Eig, Repert. Spec. Nov. Regni Veg. Beih. 55:144 (1929). TYPE: not indicated.

Aegilops ovata var. eventricosa Eig, Repert. Spec. Nov. Regni Veg. Beih. 55:144 (1929); Triticum ovatum t eventricosum (Eig) Hayek in Hayek & Markgr., Repert. Spec. Nov. Regni Veg. Beih. 30, 3:224 (1932). SYN: [1] France: Rochelle; 1858; A.-H. Letourneaux; [2] Spain: Andalusia, Malaga; 1913; M. Brandt; [3] Morocco: Bekrit, valley of Admerissen; 1925; E. Jahandiez 607; [4] Algeria: Cala Rana, Lalla Khedidja; 30 Jun 1879; H.L. Duhamel du Monceau; [5] Greece: Crete; 1928; Piner; [6] "Palestine" [Lebanon/Syria]: [Mt.] Hermon [Jabal ash-Shaykh]; 1924; A. Eig.

Aegilops neglecta Req. ex Bertol., Fl. ital. 1:787 (1834). Triticum neglectum (Req. ex Bertol.) Greuter in Greuter & Rech. f., Boissiera 13:171 (1967). HOL: BOLO; France: Avignon; 1833; E. Requien.Aegilops ovata L., Sp. pl. 2:1050 (1753), pro parte. Aegilops triaristata Willd., Sp. pl. 4:943 (1806), nom. illeg.; Triticum triaristatum (Willd.) Gren. & Godron, Fl. France 32:602 (1856); Aegilops ovata ß [var.] triaristata (Willd.) Bluff, Nees & Schauer, Comp. fl. German. 11: 209 (1836); Triticum ovatum subsp. triaristatum (Willd.) Asch. & Graebner, Syn. mitteleur. Fl. 21:703 (1901); Triticum ovatum var. triaristatum (Willd.) Stoj. & Stef., Fl. Bulg. 1:168 (1924); Aegilops ovata subsp. triaristata (Willd.) Rouy, Fl. France 14:333 (1913). TYPE: illustration in J. Scheuchzer, Agrostographia, pl. 1, f. 2A, B, C. (1719) [Greuter in Greuter & Rech. f., Boissiera 13:170 (1967)].Aegilops algeriensis Gand., Oesterr. Bot. Z. 31:82 (1881). HOL: LY (Gandoger Herb.); Algeria: near Constantine [Qacentina]; J.E. Chouette 398.Aegilops calida Gand., Oesterr. Bot. Z. 31:81 (1881). SYN: [1] LY (Gandoger Herb.); Italy: Mt. Brisighella near Faenza; Jun 1875; L. Caldesi; [2] LY; Croatia: Dalmatia, near Salona; R. de Visiani.

Aegilops campicola Gand., Oesterr. Bot. Z. 31:82 (1881). HOL: LY (Gandoger Herb.); France: Bouches-du-Rhône, Martigues; A. Autheman.

Aegilops mesantha Gand., Oesterr. Bot. Z. 31:82 (1881). HOL: LY (Gandoger Herb.); Italy: Tuscany, Florence; C.P.S. Sommier.

Aegilops viridescens Gand., Oesterr. Bot. Z. 31:81 (1881). HOL: LY (Gandoger Herb.); France: Les Arcs; C. Verriet-Litardičre.

 

Aegilops triaristata var. brachychaeta Font Quer, Treb. Inst. Catalana Hist. Nat. 5:54 (1924). HOL: BC; Spain: Gamoral, near Burgos; P. Font Quer 235.

 

Aegilops triaristata subsp. contorta Zhuk., Trudy Prikl. Bot. 18:479 (1928); Aegilops neglecta var. contorta (Zhuk.) K. Hammer, Feddes Repert. 91:240 (1980). LEC: WIR 1154; "Asia Minor" [Turkey]: Lydia, Boz-Dagh [Boz Dag], 500 m; 20 Jun 1927; P.M. Zhukovsky [designated by Zhukovsky in WIR].

Aegilops triaristata subsp. intermixta Zhuk., Trudy Prikl. Bot. 18:479 (1928). TYPE: not designated.

Aegilops triaristata var. ochreata Zhuk., Trudy Prikl. Bot. 18:480 (1928); Aegilops neglecta var. ochreata (Zhuk.) K. Hammer, Feddes Repert. 91:240 (1980). TYPE: "Tauria" [Ukraine: Crimea [Krymskij poluostrov]]; N.A. Busch.

Aegilops triaristata var. hirtula Zhuk., Trudy Prikl. Bot. 18:480 (1928). SYN: [1] "Tauria" [Ukraine: Crimea [Krymskij poluostrov]]; [2] [Greece:] Attica [Attiki]; [3] Macedonia; [4] Turkey: European Turkey.

Aegilops triaristata subsp. recta Zhuk., Trudy Prikl. Bot. 18:478 (1928); Aegilops recta (Zhuk.) Chennav., Acta Hort. Gothob. 23:165 (1960); Triticum rectum (Zhuk.) Bowden, Canad. J. Genet. Cytol. 8:135 (1966); Aegilops neglecta subsp. recta (Zhuk.) K. Hammer, Feddes Repert. 91:240 (1980). LEC: WIR 1160; Algeria: Algiers, Kabylie, Fort National, on stony soils; 29 Jul 1926; N.I. Vavilov 246 [designated by Zhukovsky in WIR].

Aegilops triaristata var. trojana Eig, Bull. Soc. Bot. Genčve, 19:328 (1928); Aegilops neglecta var. trojana (Eig) K. Hammer, Feddes Repert. 91:240 (1980). TYPE: Turkey: Thymbra, near Troja; 1889; Calvert.
Aegilops triaristata subsp. contracta Eig, Repert. Spec. Nov. Regni Veg. Beih. 55:141 (1929); Aegilops neglecta subsp. contracta (Eig) H. Scholz, Willdenowia 19:105 (1989). TYPE: Turkey: Cilicia, Trache; Apr 1872; Péronin.

Aegilops triaristata subsp. quadriaristata Eig, Repert. Spec. Nov. Regni Veg. Beih. 55:140 (1929); Aegilops neglecta var. quardiaristata (Eig) K. Hammer, Feddes Repert. 91:240 (1980). TYPE: Turkey: Taurus; 1926, E. Baur.

Aegilops tauschii Cosson, Notes pl. crit. 69 (1850) [dated 1849]. Triticum tauschii (Cosson) Schmalh., Fl. ssrednej jushnoj Rossii 2:652 (1897); Patropyrum tauschii (Cosson) A. Löve, Biol. Zentralbl. 101:206 (1982). Triticum aegilops P. Beauv. ex Roemer & Schultes, Syst. veg. 2:769 (1817). Aegilops squarrosa sensu Tausch, Flora 20:108 (1837), non L. (1753). LEC: illustration (pl. 50, f. 1) in J.C. Buxbaum's Plantarum minus cognitarum Centuria 1 (1728) [Slageren, Wild Wheats 328 (1994)].

Aegilops squarrosa ß [var.] meyeri Griseb. in Ledeb., Fl. ross. 4:326 (1852); Aegilops squarrosa subsp. meyeri (Griseb.) Zhuk., Trudy Prikl. Bot. 18:549 (1928); Aegilops tauschii var. meyeri (Griseb.) Tzvelev, Zlaki SSSR 157 (1976). HOL: LE; Azerbaijan: near Sallian; C.A. Meyer.

Aegilops squarrosa t [var.] pubescens Regel, Trudy Imp. S.-Peterburgsk. Bot. Sada 7:577 (1881). HOL: LE: [Uzbekistan/Tajikistan:] valley of the Sarawschan [Zeravsan] River, Luz (Uluss) River to Dzhan (Djam); 20 Mar 1869; O. Fedtschenko.

Aegilops squarrosa var. brunnea Popova, Trudy Prikl. Bot. Selekts. 13:477 (1923); Aegilops tauschii forma brunnea (Popova) K. Hammer, Feddes Repert. 91:233 (1980). SYN: "Turkestan" [Uzbekistan]: Samarkand, Syr-Darja [Syrdarja]; G.M. Popova.

Aegilops squarrosa var. ferruginea Popova, Trudy Prikl. Bot. Selekts. 13:477 (1923); Aegilops tauschii forma ferruginea (Popova) K. Hammer, Feddes Repert. 91:233 (1980). SYN: "Turkestan" [Uzbekistan]: Samarkand, Syr-Darja [Syrdarja]; G.M. Popova.

Aegilops squarrosa subsp. salinum Zhuk., Trudy Prikl. Bot. 18:549 (1928); Patropyrum tauschii subsp. salinum A. Löve, Feddes Repert. 95:493 (1984). HOL: WIR 2726; Azerbaijan: near Sallian; 3 Apr 1830; R.F. Hohenacker.

Aegilops squarrosa var. anathera Eig, Repert. Spec. Nov. Regni Veg. Beih. 55:90 (1929); Aegilops tauschii var. anathera (Eig) K. Hammer, Feddes Repert. 91:234 (1980). SYN: [1] Afghanistan; 1862; W. Griffith; [2]; Georgia: Elisabethpol; 1834; [3] LE; [Russia:] Dagestan; 1898; F.H. Alexeenko; [4] G (G-BOIS) [Turkey:] Cataonian Taurus; 1865; H.K. Haussknecht.

Aegilops squarrosa var. strangulata Eig, Bull. Soc. Bot. Genčve, Ser. 2, 19:328 (1928); Aegilops squarrosa subsp. strangulata (Eig) Eig, Repert. Spec. Nov. Regni Veg. Beih. 55:90 (1929); Aegilops tauschii subsp. strangulata (Eig) Tzvelev, Novosti Sist. Vyssh. Rast. 10:37 (1973); Patropyrum tauschii subsp. strangulatum (Eig) A. Löve, Feddes Repert. 95:234 (1984); Aegilops strangulata (Eig) Tzvelev, Bot. Zhurn. Moscow & Leningrad) 78:88 (1993). SYN: [1] LE, P; "Persia" [Iran]: Gulf of Astrabad (Asterabad), Gorgan; [2] LE (Fischer Herb.); Azerbaijan: Apcheron [Apseronski] Peninsula.

Aegilops triuncialis L., Sp. pl. 2:1051 (1753). Triticum triunciale (L.) Raspail, Ann. Sci. Nat. (Paris) 5:435 (1825); Aegilopodes triuncialis (L.) A. Löve, Biol. Zentralbl. 101:207 (1982). LEC: LINN 1218.8; Spain; P. Loefling 701b [Bowden, Canad. J. Bot. 37:675 (1959)].

Aegilops squarrosa L., Sp. pl. 2:1051 (1753). LEC: LINN 1218.9; "squarrosa 3" [Bowden, Canad. J. Genet. Cytol. 8:133 (1966)].

Aegilops echinata C. Presl, Cyper. Gramin. siculae 47 (1820). HOL: PRC; [Italy: Sicily,] Messina; Jul 1817; C. Presl.

Aegilops coratica Gand., Oesterr. Bot. Z. 31:81 (1881). HOL: LY (Gandoger Herb.); Croatia: Dalmatia, Fiume [Rijeka], Martinscica; L. Rossi 6046.

Aegilops triuncialis var. breviaristata Hackel in Henry Groves [of Florence], Nuovo Giorn. Bot. Ital. 19:212 (1887). TYPE: Italy: Gallipoli; Henry ["Enrico"] Groves.

Aegilops triuncialis var. albescens Popova, Trudy Prikl. Bot. Selekts. 13:476 (1923). SYN: [1] "Persia" [Iran]; N.I. Vavilov; [2] "Turkestan" [Uzbekistan]: Syr-Darja [Syrdarja]; G.M. Popova.

Aegilops triuncialis var. brunnea Popova, Trudy Prikl. Bot. Selekts. 13:476 (1923); Aegilops triuncialis forma brunnea (Popova) K. Hammer, Feddes Repert 91:238 (1980). TYPE: "Turkestan, Turkomania" [Turkmenistan]; G.M. Popova.

Aegilops triuncialis var. flavescens Popova, Trudy Prikl. Bot. Selekts. 13:475 (1923). SYN: [1] "Persia" [Iran]: Hamadan; N.I. Vavilov; [2] "Turkestan, Turkomania" [Turkmenistan]; G.M. Popova; [3] [Uzbekistan:] Syr-Darja [Syrdarja]; G.M. Popova; [4] [Uzbekistan:] Samarkand; G.M. Popova.

Aegilops triuncialis var. nigro-albescens Popova, Trudy Prikl. Bot. Selekts. 13:476 (1923); Aegilops triuncialis forma nigro-albescens (Popova) K. Hammer, Feddes Repert 91:238 (1980). SYN: [1] "Persia" [Iran]; N.I. Vavilov; [2] [Uzbekistan:] Syr-Darja [Syrdarja]; G.M. Popova; [3] [Uzbekistan:] Samarkand; G.M. Popova.

Aegilops triuncialis var. nigro-ferruginea Popova, Trudy Prikl. Bot. Selekts. 13:476 (1923); Aegilops triuncialis forma nigro-ferruginea (Popova) K. Hammer, Feddes Repert 91:238 (1980). SYN: "Turkestan" [Uzbekistan]: Syr-Darja [Syrdarja]; G.M. Popova.

Aegilops triuncialis var. nigro-flavescens Popova, Trudy Prikl. Bot. Selekts. 13:475 (1923); Aegilops triuncialis forma nigro-flavescens (Popova) K. Hammer, Feddes Repert 91:238 (1980). SYN: [2] "Persia" [Iran]: Hamadan; N.I. Vavilov; [2] "Turkestan" [Uzbekistan]: Syr-Darja [Syrdarja]; G.M. Popova.

Aegilops triuncialis var. nigro-rubiginosa Popova, Trudy Prikl. Bot. Selekts. 13:476 (1923); Aegilops triuncialis forma nigro-rubiginosa (Popova) K. Hammer, Feddes Repert 91:238 (1980). SYN: "Turkestan" [Uzbekistan]: Syr-Darja [Syrdarja]; G.M. Popova.

Aegilops triuncialis var. rubiginosa Popova, Trudy Prikl. Bot. Selekts. 13:475 (1923); Aegilops triuncialis forma rubiginosa (Popova) K. Hammer, Feddes Repert 91:238 (1980). SYN: "Turkestan" [Uzbekistan]: Syr-Darja [Syrdarja]; G.M. Popova.

Aegilops triuncialis var. nigriaristata Flaksb., Trudy Prikl. Bot. Selekts. 13:484 (1923). TYPE: [Turkmenistan:] Transcaspica province, Aschabad district, Kopet-Dagh [Kopetdag] Mountains, Tchuli; 28 May 1914; A.K. Golbeck.

Aegilops triuncialis var. glabrispica Eig, Bull. Inst. Agric. Nat. Hist. Tel-Aviv 6:71 (1927). SYN: [1] HJU; "Palestine" [Lebanon/Syria]: Mt. Hermon [Jabal ash-Shaykh], plain of Ein-Jinneh, about 1500 m; 21 Jul 1924; A. Eig; [2] HJU; "Palestine" [Lebanon/Syria]: summit of Mt. Hermon; 18 Jun 1924; A. Eig.

Aegilops triuncialis var. pubispica Eig, Bull. Inst. Agric. Nat. Hist. Tel-Aviv 6:72 (1927). SYN: [1] HJU; "Palestine" [Lebanon/Syria]: Mt. Hermon [Jabal ash-Shaykh], plain of Ein-Jinneh, about 1500 m; 21 Jul 1924; A. Eig; [2] E, HJU; [Syria]: east of Banias [Baniyas]; 10 May 1911; F.S. Meyers 1801 and J.E. Dinsmore.

Aegilops triuncialis var. assyriaca Eig, Bull. Soc. Bot. Genčve, Ser. 2, 19:323 (1928). SYN: [1] G, JE, LE, IS, W, Z; "Assyria" [Iraq]: eastern Assyria, Kuh-Sefin Mountains; 4 Jun 1893; J.F.N. Bornmüller 1899; [2] B, JE, K; "Assyria" [Iraq]: eastern Assyria, Kuh-Sefin Mountains, Kerkuk; 26 Apr 1893; J.F.N. Bornmüller 1900; [3] B. JE, K; "Assyria" [Iraq]: eastern Assyria, Kuh-Sefin Mountains, Kerkuk [Kirkuk]; 26 Apr 1893 J.F.N. Bornmüller 1903.

Aegilops triuncialis subsp. caput-medusae Zhuk., Trudy Prikl. Bot. 18:499 (1928). LEC: WIR 1328; "Asia Minor" [Turkey]: Lydia, Boz-Dagh [Boz Dag]; P.M. Zhukovsky [Slageren, Wild Wheats 349 (1994)].

Aegilops triuncialis subsp. fascicularis Zhuk., Trudy Prikl. Bot. 18:499 (1928). LEC: WIR 1372; Spain; N.I. Vavilov [Slageren, Wild Wheats 349 (1994)].

Aegilops triuncialis var. hirta Zhuk., Trudy Prikl. Bot. 18:500 (1928). TYPE: not designated.

Aegilops triuncialis var. muricata Zhuk., Trudy Prikl. Bot. 18:500 (1928). TYPE: not designated.

Aegilops triuncialis var. constantinopolitana Eig, Repert. Spec. Nov. Regni Veg. Beih. 55:133 (1929). HOL: G (G-BOIS); [Turkey:] Constantinople [Istanbul], Chalki Island; 1876; Murmann.

Aegilops ventricosa Tausch, Flora 39:108 (1837). Triticum ventricosum (Tausch) Ces., Pass. & Gibelli, Comp. fl. ital. 14:86 (1869); Gastropyrum ventricosum (Tausch) A. Löve, Biol. Zentralbl. 101:208 (1982). NEO: G; [Spain: Granada,] Sierra Nevada; P.E. Boissier [Slageren, Wild Wheats 392 (1994)].

Aegilops fragilis Parl., Fl. ital. 1:515 (1850); Triticum fragile (Parl.) Ces., Pass. & Gibelli, Comp. fl. ital. 14:87 (1869), non Roth (1800); Aegilops ventricosa ß [var.] fragile (Parl.) Fiori & Paol., Fl. Italia 1:109 (1896); Aegilops ventricosa subsp. fragilis (Parl.) Zhuk., Trudy Prikl. Bot. 18:523 (1928). SYN: [1] FI, PAV; [Italy: Puglia]; Jun 1848; G. Gasparrini; [2] FI, TO; [Italy: Sardinia]; G.G. Moris.

Aegilops squarrosa ß [var.] comosa Kralik in Cosson, Notes pl. crit., Ser. 1, 2[bis]:68 (1850); Aegilops ventricosa forma comosa ("Cosson" [error for Kralik]) Batt. & Trabut, Fl. Algérie 12:242 (1895); Aegilops ventricosa subsp. comosa ("Cosson" [error for Kralik]) Zhuk., Trudy Prikl. Bot. 18:522 (1928); Aegilops ventricosa var. comosa ("Cosson" [error for Kralik]), Eig, Repert. Spec. Nov. Regni Veg. Beih. 55:97 (1929). HOL: P; Algeria: near Oran; M.C. Durieu de Maisonneuve.

Aegilops squarrosa t [var.] truncata Kralik in Cosson, Notes pl. crit., Ser. 1, 2[bis]:68 (1850); Aegilops ventricosa forma truncata ("Cosson" [error for Kralik]) Batt. & Trabut, Fl. Algérie 12:242 (1895); Aegilops ventricosa subsp. truncata ("Cosson" [error for Kralik]) Zhuk., Trudy Prikl. Bot. 18:523 (1928); Aegilops ventricosa var. truncata ("Cosson" [error for Kralik]), Eig, Repert. Spec. Nov. Regni Veg. Beih. 55:97 (1929). SYN: [1] P; Algeria: near Oran; M.C. Durieu de Maisonneuve; [2] P; Algeria: near Ghelma; J.P. Kremer.

Aegilops subulata Pomel, Nouv. mat. fl. atl. 388 (1875); Triticum subulatum (Pomel) T. Durand & Schinz, Consp. fl. afric. 5:939 (1894), non Schreber in Schweigger & Körte (1811); Aegilops ventricosa var. subulata (Pomel) Maire & Weiller in Maire, Flore de l'Afrique du Nord 3:355 (1955). HOL: AL; Algeria: grassy places in mountains, Aflou; 6 Jun 1860; A.N. Pomel 171.

Aegilops ventricosa var. prostrata Sennen & Mauricio in Sennen, Bol. Soc. Ibér. Ci. Nat. 28:174 (1929). HOL: BC; Morocco: Beni-Said, Pont du Kert; Frčre Sennen 9622 and Frčre Mauricio.

Aegilops ventricosa var. obscura Miczynski, Bull. Soc. Bot. France 76:715 (1929). HOL: AL; Algeria: Berteaux (Constantine) [Qacentina]; L.O. Ducellier.
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EXCLUDED NAMES
Aegilops vagans Jordan & Fourr., Brev. pl. nov. 2:130 (1868); Triticum vagans (Jordan & Fourr.) Greuter, Boissiera 13:170 (1967). TYPE: France: southern France, near Montpellier, etc. [Slageren, Wild Wheats 244 (1994)].