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Contents: Introduction; Key; Taxa; Comment; Anatherostipa Links: Stipeae home; Intermountain Herbarium These notes prepared by M. Barkworth

Anatherstipa was the name given by Peñailillo to a group of grasses identified by Parodi (1950) as the Obtusae group. Dr. M.A. Torres proposed the name Anatherostipa for the same group in May, 1997, and her article was received by many major herbaria at the endo of that month.. Peñailillo's article, which came out in Gayana, did not reach other herbaria until August, but priority dates from when two copies are made available to appropriate institutions The editor of Gayan stated that two copies of the issue of Gayana had been made available by the publication date on the issue's cover, December, 1996, so Anatherostipa has priority over Anatherostipa.

The information on this page comes from Torres because Peñailillo merely made the necessary new combinations. Because my Spanish and Latin are limited, those more than slightly interested in the genus should be very sure to consult Torres' article.

The type of Anatherostipa is A. saltensis (Kuntze) Peñailillo. The species of the genus grow at 3,200-5,000 m in arid desert regions of the mountains in southern Bolivia and Peru, northwestern Argentina, and northern Chile. Parodi (1950) mentioned that two of the species, A. bomanii and A. saltensis [both cited as species of Stipa] were known for their toxicity. The following is a translation of Dr. Torres' key distinguishing Anatherostipa from other Stipoid genera that occur in northeastern Argentina. The generic description follows the key.


1. Mature lemmas with strongly overlapping margins that completely conceal the palea and the caryopsis; palea without veins, 1/3-1/4 the length of the lemma. Flowers either cleistogamous with 1 minute anther, much shorter than those of their chasmogamous counterparts and, normally, with 2 staminodes (these also in the chasmogamous flowers). Lemma epidermes composed of very short, silicified fundamental cells with sinuous or no sidewalls; silica cells (= idioblasts) absent or rare Nassella
1. Mature lemmas with slightly overlapping or open margins., not completely concealing the palea and caryopsis; palea 2-veined, concave or flat, either 1/2-2/3 the length of the lemma or equaling it, rarely shorter. Flowers not or only slightly differentiated into two types by the size of the anthers, the dorsal anther slightly the smaller, staminodes generally absents. Lemma epidermes not as above
2. Prophyllum 2-awned; paleal veins terminating at or near the apex. Lemma epidermes with long fundamental cells with very sinuous side walls; silica cells present or absent Anatherostipa
2. Prophyllum entire, 2-toothed, or emarginate, never 2-awned; paleal veins not extending to the apex. Lemma epidermes with short or long fundamental cells with sinuous sides walls; silica cells present, in clusters or scattered Stipa [Peñailillo has placed these species in a greatly enlarged Jarava]

Anatherostipa (Hack. ex Kuntze) Peñailillo

Plants perennial, caespitose, 10-50 cm tall, golden, ? or stramineous to ochroleucous (often conspicuously so at maturity); innovations intravaginal, prophyllum 2-awned, 2-5 cm long. Culms erect. Leaf sheaths glabrous; internodes shorter than the sheaths; auricles absent. Ligules membranous, 3-10 mm (or shorter). Leaf blades setaceous-filiform, convolute, erect or curved, 3-17 cm long, 0.3-0.5 mm in diameter, glabrous. Inflorescence very narrow, with few spikelets, exserted, 3-15 cm long; branches appressed to the rachis. Glumes 2, papery, obtuse of somewhat acute, equal or subequal, (1)-3-veined, the veins evident. Lemma fusiform, the apex only slightly attenuate, 2.5-4 mm long, the lemma body glabrous or pubescent, scabrous, shiny or slightly shiny, the lemma margins open (back of the palea visible, not completely concealed). Crown absent. Awn central, capillary, straight, arcuate, or flexuous, or reduced to a diminutive mucro, glabrous, deciduous or persistent. Callous obtuse, about 0.5 mm long, pilose with short hairs from near the base. Palea papery, flat or keeled, glabrous or pilose, as long as the body of the lemma, 2-veined, the veins extending to, or almost to, the tip. Lodicules 2-3, hyaline. Androecium of 3 stamens, the anther sacs glabrous, rarely subglabrous. Caryopsis ellipsoid, the hilum linear, straight, ventral.

Lemma epidermis composed of long fundamental cells with markedly sinuous walls. Hooks, sometimes associated with unicellular hairs often present, both of these kinds of cella having swollen bases. Silica cells, cells having a siliceous body in their interior, are sometimes present.


Anatherostipa subg. Anatherostipa Stipa sect. Anatherostipa Hackel; Stipa subg. Anatherostipa (Hackel) Speg.
Anatherostipa bomanii (Hauman) Peñailillo Stipa bomanii Hauman Argentina (Jujuy)
Anatherostipa brevis (Torres) Peñailillo Argentina (Jujuy); probably Bolivia (fide M.A. Torres)
Anatherostipa coroi F. Rojas
Anatherostipa hans-meyeri (Pilg.) F. Rojas
Anatherostipa henrardiana (Parodi) Peñailillo Stipa henrardiana Parodi Argentina (Jujuy)
Anatherostipa mucronata (Griseb.) F. Rojas Piptochaetium mucronatum Griseb.; Orysopsis mucronata (Griseb.) Parodi; Stipa saltensis Kuntze Argentina (Jujuy, Salta, Tucuman)
Anatherostipa obtusa (Nees & Meyen) Peñailillo Piptatherum obtusum Nees & Meyen; Urachne obtusa (Nees & Meyen) Trinius; Stipa obtusa (Nees & Meyen) Hitchc.; Stipa boliviensis Hackel; Oryzopzis neesii Pilger; Stipa jujuyensis Speg. Argentina (Jujuy); Bolivia; Peru
Anatherostipa orurensis F. Rojas
Anatherostipa rigidiseta (Pilger) Peñailillo Oryzopsis rigidiseta Pilger; Stipa rigidiseta (Pilger) Hitchc. Chile (Tarapacá); Peru (Azángaro); probably Bolivia (fide M.A. Torres)
Anatherostipa rosea (Hitchc.) Peñailillo
Anathertostipa saltensis (Kuntze) Peñailillo
Anatherostipa venusta (Phil.) Peñailillo Stipa venusta Philippi Chile (Antofagasta, Taparacá)