Elymus
Being compiled by M.E. Barkworth, to whom corrections and suggestions should be sent.]; Topics: Circumscription, distribution, size, cytogenetics, list of species
Circumscription: Elymus is interpreted here as comprising primarily caespitose, short-anthered species with non-disarticulating rachises, and lanceolate, somewhat flexible, glumes and their genetically close relatives. The last phrase brings into Elymus a few rhizomatous, long-anthered species (e.g., Elymus repens, E. lanceolatus), some species with almost no glumes (e.g., E. hystrix), and some with disarticulating rachises as well as narrow, awn-like glumes (e.g., E. elymoides).
The number of spikelets at each node is not important; it may be 1 (as in sect. Roegneria and many of the species that used to be placed in Agropyron) or more than 1 (as in E. viriginicus, E. canadensis, etc.).
For different viewpoints on the circumscription of Elymus, see Assadi and Runemark (1995) (who interpret it to include Thinopyrum and Pseudoroegneria plus many other segregates) or Baum et al. (1991) (who treat Roegneria and Elytrigia as distinct genera). For a revision of the South American taxa, see Seberg and Petersen (1998).
Distribution: Elymus is a cool-temperate genus. It is most abundant in Asia and North America, but extends into Europe, South America, and Australia.
Size: Elymus probably consists of around 200 species. Estimating its size is difficult because its greatest diversity is in Asia, many parts of which are poorly known botanically speaking. An additional complication is a consequence of the differing species interpretations that have been adopted. North American species are thought to have low barriers to interspecific hybridization. Interestingly, this does not seem to be true of Asian species (von Bothmer, pers. comm).
Genomic composition: Elymus is an alloploid genus, all of whose members (so far as is known) possess the St genome in combination with at least one other genome. Most species are tetraploids, but there are many hexaploids and possible some octoploids. All native North American species examined are tetraploids in which the second genome is the H genome. Many Asian species are also tetraploids, but with the Y genome as the additional genome. Other genomes known to occur in Elymus are the P and E genomes.
Table of species: The following table will be built up very slowly (unless funding becomes available). The synonyms listed are those that seem to be in recent or current use and are provided to help in interpretation of the treatment reflected by the first column. Genomic information is provided ONLY if it has been obtained via investigation of the species involved. Reports that do not list voucher specimens made at the time of the genomic investigation are asterisked. They should be repeated, with vouchers being deposited in a herbarium that regularly makes loans available. The species are listed alphabetically for now. The links below will aid in locating names in the first column.
| E. abolinii var. pluriflorus D.F. Cui | China (Xinjiang) | |||
| E. alaskanus | ||||
| E. alaskanus subsp. islandicus (Melderis) A. Love & D. Love | ||||
| E. alaskanus subsp. subalpinus (Neuman) A. Love & D. Love | ||||
| Elymus alopex B. Salomon | Iceland | |||
| E. altaicus D.F. Cui | China (Xinjiang) | |||
| E. angulatus J. Presl in C. Presl | Agropyron breviaristus Hitchc. | Peru south to Tierra del Fuego, east to western Argentina and Bolivia | 2n=28 | |
| E. anthosachnoides (Keng) A. Löve | R. anthosachnoides Keng | Southwestern China (Sichuan, Yunnan). [Lu 1993] | StY | |
| E. cacuminus B.-R. Lu & B. Salomon | Western China (Sichuan), Nepal, Sikkim, Tibet. [Lu 1993] | StY | ||
| E. canaliculatus (Nevski) Tzvelev | Agropyron canaliculatum Nevski; Roegneria canaliculata (Nevski) Ohwi; Elymus longearistatus subsp. canaliculatus (Nevski) Tzvelev; A. longearistatum var. aitchisonii Boissier | Alai, Iran, Kashmir, Pakistan (Gilgit, NWFP), Pamir, Tadzhikistan, Tibet. [Lu 1993] | StY | |
| E. caninus (L.) L. | ||||
| E. caninus var. behmii (Melderis) Jaaska | ||||
| E. caninus (L.) L.var. caninus | ||||
| E. caninus var. donianus (F.B. Whie) Jaaska | ||||
| E. caninus var. muticus (Holm.) T. Karlsson | ||||
| E. caucasicus (C. Koch) Tzvelev | Roegneria caucasicus C. Koch; Agropyron caucasicum (C. Koch) Grossh.; Agropyron roegneri (Griseb.) Boissier; R. linczevskii Czopan. | Daghestan, Turkmenia, northern Iran. [Lu 1993] | StY | |
| E. ciliaris (Trin.) Tzvelev | StY | |||
| E. cordilleranus Davidse & R.W. Pohl | Costa Rica, south through the Andes to Peru and Bolivia | 2n=28 | ||
| E. elymoides (Raf.) Swezey | Sitanion hystrix | Western North America | StH | |
| E. elymoides subsp. brevifolius (J.G. Smith) Barkworth | Sitanion brevifolius J.G. Smith | Western North America | StH | |
| E. elymoides subsp. californicus | Sitanion californicum J.G. Smith | Western North America | StH | |
| E. elymoides subsp. elymoides | Western North America | StH | ||
| E. elymoides subsp. hordeoides | Sitanion hordeoides Suksdorf | Western North America; rare | StH | |
| E. dahuricus Turcz. ex Griseb. | StYH | |||
| E. fibrosus (Schrenk) Tzvelev | ||||
| E. formosanus (Honda) A. Löve | Agropyron formosanum Honda; Roegneria formosana (Honda) Ohwi; R. formosana var. longearistata Keng; R. formosana var. pubigera Keng | Formosa Island. [Lu 1993]] | ? | |
| E. glaberrimus var. breviaristatus D.F. Cui | China (Xinjiang) | |||
| E. glaucescens Seberg | Agropyron pubiflorum (Steudel) Parodi; A. antarcticum Parodi | Southern Chile and Argentina | 2n=28 | |
| E. gmelinii (Ledeb.) Tzvelev | Central and East Asia, | StY | ||
| E. humidus (Ohwi & Sakamoto) Osada | StYH | |||
| E. jacquemontii (Hooker f.) Tzvelev | Agropyron jacquemontii Hooker f; Roegneria jacquemontii (Hooker f.) Nevski; Anthosachne jacquemontii (Hooker f.) Nevski | Western Tibet and the Himalayas, above 3900 m. [Lu 1993] | StY | |
| E. kaschgaricus D.F. Cui | China (Xinjiang) | |||
| E. longearistatus (Boissier) Tzvelev | Agropyron longearistatum (Boissier) Boissier; Anthosachne longearistata (Boissier) Nevski; Roegneria longearistata (Boissier) Drobov; A. longearistata var. haussknechtii Boissier | Gissar-Darvaz, Iran, Tadzhikistan. [Lu 1993] | StY | |
| E. magnicaespes D.F. Cui | China (Xinjiang) | |||
| E. mendocinus (Parodi) A. Love | Agropyron mendocinum Parodi | Argentina (endemic around Mendoza) | 2n=56 | |
| E. mutabilis Drobov) Tzvelev | ||||
| E. mutabilis var. nemoralis S.L. Chen ex D.F. Cui | China (Xinjiang) | |||
| E. nipponicus Jaaska | StY | |||
| E. parodii Seberg & G. Petersen | A. condensatum J. Presl | Southern Brazil, Uruguay, and northern Argentina | 2n=28 | |
| E. patagonicus Speg. | Southern Argentina and Tierra del Fuegao | 2n-42 | ||
| E. repens (L.) Gould | Cosmopolitan weedy species | |||
| Elytrigia repens subsp. longearistata N.R. Cui | China (Xinjiang) | |||
| Elytrigia repens var. subulatum Seberg & G. Petersen | ||||
| E. retroflexus B.-R. Lu & B. Salomon | Tibet [Lu 1993] | StY | ||
| E. x russellii (Melderis) Cope | Agropyron russellii Melderis | Pakistan (near Makerum, Hispar, and Biafo glaciers). [Lu 1993] | ? | |
| E. scabriglumis (Hackel) A. Love | Argentina and Chile | 2n=28 | ||
| E. schugnanicus (Nevski) Tzvelev | Agropyron schuganicum Nevski; Roegneria schugnanica (Nevski) Nevski | Gissar-Darvaz, Pamir and, reportedly, northeastern Iran andwestern Himalayas. [Lu 1993] | ? | |
| E. semicostatus (Steud.) Melderis | Triticum semicostatum (Nees ex Steud.; Roegneria semicostata (Sted.) Kitag. | StY (Svitashev et al. 1998) | ||
| E. serotinous (Keng) A.Löve | R. serotina Keng | Central China (Shaanxi). [Lu 1993] | ? | |
| Elymus sibiricus L. | ||||
| E. sinkiangensis D.F. Cui | China (Xinjiang) | |||
| E. stebbinsii Gould | Agropyron parishii Scribner J.G. Smith | California (See Barkworth 1997) | ||
| E. stebbinsii subsp. septentrionalis Barkworth | California (See Barkworth 1997) | |||
| E. tschimganicus var. glabrispiculus D.F. Cui | China (Xinjiang) | |||
| E. tsukushiensis Honda | StYH | |||
| Elymus wawawaiensis J.R. Carlson & Barkworth | StH | |||
| E. yangii B.-R. Lu | Western China and Tibet, between 3-4200 m [Lu 1993] | StY | ||
| E. yubaridakensis (Honda) Ohwi | Japan | |||
| Elymus antiquus (Nevski) Tzvelev | Agropyron antiquus Nevski; A. microlepis Bor; Roegneria parvigluma Keng | Western and northwestern China (Qinghai, Sichuan, Yunnan), Nepal, and Tibet. [Lu 1993] | StY | |
| Elymus brevipes (Keng) A. Löve | Roegneria brevipes Keng | Western and northwestern China (Gansu, Qinghai, Sichuan), and Tibet. [Lu 1993] | StY | |
| Elymus burchan-buddae (Nevski) Tzvelev) | Agropyron burchan-buddae Nevski; A. nutans Keng; A. breviglume Keng, Roegneria breviglumis (Keng) Keng | Western and northwestern China (Gansu, Qinghai, Sichuan, Xinjiang, Yunnan), and Tibet. Probably also in Nepal and northern India. [Lu 1993] | StY | |
| Elymus sclerus A. Löve | Roegneria dura (Keng) Keng | Western and northwestern China (Gansu, Qinghai, Sichuan, Xinjiang), and Tibet. [[Lu 1993] | StY | |
| E. alaskanus (Scribner & Merr.) A. Löve | ||||
| E. alaskanus (Scribner &Merr.) A. Löve subsp. alaskanus | Agropyron boreale (Turcz.) Drobow; Roegneria borealis (Turcz.) Nevski Elymus kronokensis subsp. borealis (Turcz.) Tzvelev | Alaska, Canada (Yukon, Northwest Territories) | ||
| E. alaskanus subsp. latiglumis (Scribner & J.G. Sm.) A. Löve | Agropyron latiglume (Scribner & J.G. Sm.) Rydb.; A. caninum var. latiglume (Scribner & J.G. Sm.) Pease & A.H. Moore; Elymus trachycaulus var. latiglume (Scribner & J.G. Sm.) Beetle; A. violaceum (Hornem.) Lange; R. violacea (Hornem.) Melderis | Canada (Alberta, BC) USA (Alaska, Idaho, Washington, Montana, Wyoming) | ||
| E. hyperarcticus (Polunin) Tzvelev | Agropyron boreale var. hyperarcticum (Polunin) A. Löve & D. Löve; Roegneria borealis subsp. hyperarctica (Polunin) A. Löve & D. Löve; Elymus hyperarcticus (Polunin) Tzvelev; E. sajanensis subsp. hyperarcticus (Polunin) Tzvelev; E. alaskanus subsp. hyperarcticus (Polunin) A. Löve & D. Löve | USA (Alaska); Russia (Arctic) | ||
| E. californicus (Bolander ex Thurber) Gould | Hystrix californica Bolander ex Thurber [Molecular data suggest that it belongs in Leymus] | USA (California, around San Francisco Bay). | 56 | |
| E. transhyrcanus (Nevski) Tzvelev | Roegneria transhyranicus Nevski; Agropyron transhyranicum (Nevski) Bondartseva; Roegneria leptoura Nevski; Semiostachys leptoura (Nevski) Drobov; Elytrigia vvedenskyi Drobov | Iran and central Asia | StStH | |
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