| Mary E. Barkworth |
Plants perennial; monoecious,
staminate and pistillate spikelets evidently distinct, located in
the same inflorescences, pistillate spikelets below the staminate
spikelets. Culms 0.7-5 m. Leaves
not aromatic; sheaths open; ligules membranous, erose to
ciliate.
Inflorescences terminal and axillary, panicles of 1-several subdigitate
to racemose rames; rames with pistillate spikelets proximally
and staminate spikelets distally; disarticulation in the rames,
beneath the pistillate spikelets and at the base of the staminate portions. Pistillate
spikelets
exposed, solitary, embedded in the indurate rame axes; lower glumes coriaceous,
closing the hollows in the rachises and concealing the florets; upper
glumes
similar but smaller; lower florets sterile; upper florets pistillate;
lemmas and paleas hyaline, unawned; styles 2, not
fused.
Staminate spikelets paired, both sessile or both subsessile, or
1 sessile and the other pedicellate; glumes coriaceous, chartaceous,
or membranous;
lemmas and paleas hyaline, unawned. Pedicels (when
present) not fused to the rame axes. x = 9. The origin of the
name is unknown.
Tripsacum is a genus of 12 species, all of which are native to tropical
and subtropical regions of the Western Hemisphere; three are native to the Flora
region. They are good forage grasses, but are rarely sufficiently abundant in
the Flora region to be important in this regard. The genus is of interest
to plant breeders because of its relationship to Zea.
Measurements of the pistillate spikelets are based on measurements
of the lower glumes of the sessile spikelets, the remainder of the spikelet
being concealed between the rachis and the lower glume.
1 |
Staminate spikelets in sessile-pedicellate pairs,
the pedicels almost flat to plano-convex in cross section, 2-5 mm long,
less than 0.3 mm wide; glumes usually membranous (sect. Fasciculata)
..... 1. T. lanceolatum |
Staminate spikelets sessile, subsessile,
or in sessile-pedicellate pairs, the pedicels triangular in
cross section, to 2 mm long and about 0.5-0.8 mm wide; glumes
somewhat coriaceous (sect. Tripsacum)
(2) |
|
Blades 9-35(45) mm wide, flat; culms 1-2(4)
m tall ..... 2. T. dactyloides |
|
Blades 1-7(15) mm wide, involute or folded; culms to 1 m
tall ..... 3. T. floridanum |
Tripsacum sect. Fasciculata Hitchc.
Staminate spikelets in sessile-pedicellate pairs, the
pedicels slender and relatively flexible.
1. Tripsacum lanceolatum Rupr. ex E. Fourn.
Mexican Gamagrass
Plants rhizomatous. Culms 1-2 m tall, 2-4 mm thick. Lower sheaths
hispid; upper sheaths essentially glabrous; ligules erose, not ciliate;
blades to 100 cm long, 8-30 mm wide, glabrous or slightly pubescent. Terminal
inflorescences with 4-7(10) rames. Pistillate spikelets 2-3 mm wide,
beadlike in appearance. Staminate spikelets in sessile-pedicellate pairs;
glumes 5-10 mm long, 1.5-2 mm wide, usually membranous, acute; pedicels
2-5 mm long, less than 0.3 mm wide, almost flat to plano-convex in cross section,
flexible. 2n = 72.
Tripsacum lanceolatum grows in moist soil (often in canyon bottoms)
of mountains from southeastern Arizona and southwestern New Mexico through
Mexico to Guatemala. It has not been found in New Mexico since the 1800s.
Tripsacum L. sect. Tripsacum
Staminate spikelets in sessile pairs or 1 member of
each pair on a short, stout pedicel.
2. Tripsacum dactyloides (L.) L.
Eastern Gamagrass
Plants with short, knotty rhizomes. Culms 1-2(4) m tall, 3-5 mm
thick, clumped. Sheaths usually glabrous, occasionally slightly pilose;
ligules ciliate; blades 30-75(120) cm long, 9-35(45) mm wide,
flat, usually glabrous, tapering to attenuate apices. Terminal inflorescences
erect, with (1)2-3(6) rames; rames 12-25 cm. Pistillate spikelets
6-8 mm long, 3-5.5 mm wide. Staminate spikelets all sessile or subsessile;
glumes 5-12 mm, coriaceous, blunt, acute, or bifid; pedicels,
when present, about 1 mm long, 0.5-0.8 mm wide, triangular in cross section,
rigid. 2n = 36, 54, 72.
Tripsacum dactyloides grows in water courses and limestone outcrops
from the central and eastern United States through Mexico to northern
South America. Plants from the United States and northern Mexico belong
to Tripsacum dactyloides (L.) L. var. dactyloides. They
differ from those of the other two varieties in their erect stems and
sessile staminate spikelets. Narrow-bladed plants of T. dactyloides from
Texas resemble
T. floridanum, but on transplanting
to favorable conditions develop the wider blades characteristic of T.
dactyloides.
The two species can hybridize; the hybrids are partially sterile.
Growing Tripsacum dactyloides for forage has proven
practical only in South America. It is also used as an ornamental grass, the
chief attraction being its foliage.
3. Tripsacum floridanum Porter ex Vasey
Florida Gamagrass
Plants with short, thick rhizomes. Culms to 1 m tall, to 2 mm thick,
usually solitary or in small clumps. Sheaths glabrous; blades to
60 cm long, 1-7(15) mm wide, involute or folded, glabrous. Terminal inflorescences
erect, with 1-2 rames. Pistillate spikelets 3.5-4.5 mm wide. Staminate
spikelets sessile-pedicellate; spikelets 5-7 mm; glumes coriaceous,
acute; pedicels to 2 mm long, to 0.5 mm wide, triangular in cross section.
2n = 36.
Tripsacum floridanum grows along roadsides and in pine woods, often in
wet soils, of Florida and Cuba. It is grown as an ornamental, but it reseeds rather
too readily under some conditions. Reports of T. floridanum from Texas
are based on narrow-bladed specimens of T.
dactyloides.