| Gerrit Davidse |
Plants perennial (rarely annual);
cespitose. Culms 0.5-8 m, weakly lignified, erect, arching, climbing,
or decumbent, rooting at the nodes. Sheaths open; ligules membranous,
sometimes ciliate; pseudopetioles sometimes present; blades linear
to ovate, bases slightly to strongly asymmetric. Inflorescences open
or contracted panicles, rachises usually visible, even distally, spikelets attached
obliquely to the pedicels; disarticulation below the glumes. Spikelets
subglobose to globose, with 2 florets. Glumes membranous, apices
lanate pubescent, abruptly apiculate; lower glumes 1/3-2/3 as long as
the spikelets, 5-13-veined, bases saccate, margins overlapping; upper glumes
about as long as the upper florets, not saccate, 7-15-veined; lower florets
sterile or staminate; lower lemmas membranous, apices lanate pubescent,
abruptly apiculate; lower paleas present, sometimes reduced; upper
florets stipitate, bisexual, appearing to be mucronate or acuminate; upper
lemmas indurate, usually broadly elliptic to obovate, margins enclosing
the edges of the paleas, apices obtuse, somewhat woolly pubescent, usually dark
brown at maturity; upper paleas similar to the lemmas, but saccate below
and gibbous above. Caryopses plano-convex, ovoid, or nearly orbicular;
embryo about 1/2 as long as the caryopses; hila oblong to nearly
round. x = 9. Name from the Greek lasios, woolly, and akis,
point, referring to the tuft of wool at the apex of the spikelets.
Lasiacis is a neotropical genus of 16 species that extends from southern
Florida to Peru and Argentina. Two species are native to the Flora region.
The shiny black color of its mature florets and the oil-filled cells of the
inner epidermes of the glumes and sterile lemmas distinguish Lasiacis
from all other grasses. Birds are a common dispersal agent.
In Lasiacis the upper florets appear to be mucronate
or acuminate. The mucro or acuminate apex is formed by the tuft of hairs at
the apex of the upper floret.
1 |
Leaf blades linear-lanceolate to narrowly
lanceolate, 3-16 cm long, 3-30 mm wide ..... 1. L.
divaricata |
Leaf blades ovate to broadly lanceolate, 2-16 cm long, 8-56
mm wide ..... 2. L. ruscifolia |
1. Lasiacis divaricata (L.) Hitchc.
Plants perennial; cespitose. Culms (0.5)1-5(7)
m long, arching, clambering over brush or bending to the ground if unsupported.
Sheaths mostly glabrous, margins and throats ciliate; ligules
0.8-13 mm, glabrous or ciliate; blades (3)5-12(16) cm long, 3-20(30)
mm wide, linear-lanceolate to narrowly lanceolate, lower blades often deciduous.
Panicles 2-12(20) cm; branches to 12 cm, with widely-spaced spikelets;
lower branches usually reflexed; upper branches widely spreading.
Spikelets 3.5-4.5 mm, obovate. Lower glumes 1.2-2.5 mm; lower
florets sterile; upper florets 3.4-4 mm long, 1.9-2.4 mm wide, whitish
to brown at maturity; anthers about 2 mm, white; stigmas purple.
Caryopses whitish. 2n = 36.
Lasiacis divaricata is a Caribbean species. Its range extends from Florida
through the West Indies to Mexico, Panama, and northern Venezuela. In Florida,
it usually grows in hammocks, but occasionally in pinelands. The whitish to
brown upper florets are unusual in the genus.
Plants in the Flora region belong to Lasiacis divaricata
var. divaricata, which differs from the other two varieties in having
panicles with fewer spikelets and panicle branches that are usually reflexed.
2. Lasiacis ruscifolia (Kunth) Hitchc.
Plants perennial; cespitose. Culms 1-8 m long, 5-12 mm thick, hollow,
arching or clambering. Sheaths glabrous, puberulent, or hispid with papillose-based
hairs, margins ciliate; ligules 0.2-1 mm, glabrous or ciliate; blades
2-16 cm long, 8-56mm wide, ovate to ovate-lanceolate. Panicles 2-22 cm;
lower branches to 9 cm, divergent. Spikelets 2.6-4 mm, globose;
lower glumes 1.2-2.2 mm, 9-13-veined; lower florets sterile; upper
glumes 11-13-veined; upper florets 2.8-3.6 mm long, 2-2.9 mm wide,
dark brown to grayish-black at maturity; upper lemmas usually with a distinct
shelf at the base, from which a sterile projection often arises; upper paleas
usually deeply concave; anthers 1.4-2.3 mm, white; stigmas white.
Caryopses 2-2.5 mm. 2n = 36.
The range of Lasiacis ruscifolia extends from southern Florida to Peru.
Plants in the Flora region belong to Lasiacis ruscifolia (Kunth)
Hitchc. var. ruscifolia, which differs from L. ruscifolia var. velutina
(Swallen) Davidse in being scabrous or puberulent, rather than velutinous, on
the panicle branches.