| Mary E. Barkworth |
Plants perennial. Culms 50-350
cm, decumbent, often rooting from the lower nodes; nodes glabrous; internodes
glabrous, filled with spongy aerenchyma. Leaves evenly distributed; ligules
membranous. Inflorescences narrow, condensed, cylindrical or spikelike
panicles; branches appressed but not fused to the rachises, densely and
evenly short ciliate on the ridges. Spikelets dorsally compressed, narrowly
lanceolate, with 2 florets. Glumes unequal, membranous, with a definite
rachilla internode between the 2 glumes; lower glumes no more than 1/2
as long as the spikelet, 1-3-veined; upper glumes subequal to or longer
than the lower lemmas, membranous, not saccate, 3-7-veined; lower florets
sterile; lower lemmas similar to the upper glumes; upper florets
bisexual, much shorter than the upper glumes and lower lemmas; upper lemmas
membranous to leathery, white at maturity, margins clasping the edges of the
paleas basally but not distally, apices acute; paleas similar in texture
to the lemmas; lodicules 2; anthers 3; styles 2, fused. Caryopses
falling free from the lemmas and paleas. x = 10. Name from the Greek
hymen, membrane, and achne, chaff, alluding to the membranous upper
lemma, an unusual texture for the upper lemma in members of the Paniceae.
Hymenachne is a pantropical genus of approximately five species. It is
unusual in the Paniceae in that all its members grow in aquatic or swampy
habitats. It differs from Sacciolepis in
its aerenchyma-filled internodes, a character that is probably an adaptation to
its aquatic habitat. One species is native to the Flora region.
1. Hymenachne amplexicaulis (Rudge) Nees
West Indian Marsh Grass
Plants perennial. Culms to 3.5 m tall, 1 cm or more
thick, decumbent. Ligules 1-2.5 mm, brownish; blades 15-33 cm
long, 12-28 mm wide, lax, flat, glabrous. Panicles 10-40 cm long, 0.7-1.2
cm thick, spikelike, dense, sometimes lobed near the base; basal branches
1.5-5 cm, strictly erect. Spikelets 3.5-5 mm, lanceolate, acuminate.
Lower glumes 1-1.7 mm, 3-veined; upper glumes 2.8-3.9 mm; lower
lemmas 3.6-4.6 mm, longer than the upper glumes, attenuate to subaristate;
lower paleas absent; upper lemmas 2.5-3.5 mm; anthers 1.1-1.2
mm. 2n = 10.
In the Flora region, Hymenachne amplexicaulis is known only from
low, wet pastures in southern Florida and it is rare even in that state. It
is more abundant in the remainder of its range which extends through Mexico
to Argentina.