| John W. Thieret† |
Plants perennial; cespitose,
sometimes stoloniferous. Culms 10-70 cm, herbaceous, erect
to decumbent, basal branching extravaginal. Leaves not aromatic; sheaths open; auricles
absent; ligules membranous, truncate; blades flaccid, linear
to lanceolate. Inflorescences terminal, of long-pedunculate, solitary,
1-sided rames (axillary rames occasionally present) with more than 1 spikelet
unit; rame
internodes clavate, glabrous; disarticulation in the rames. Spikelets
in heteromorphic sessile-pedicellate pairs, pedicellate spikelets absent
or rudimentary. Sessile spikelets imbricate, not embedded in the
rame axes, dorsally compressed, with 2 florets, unawned; calluses truncate; glumes exceeding
the florets, differing in shape; lower glumes 4-9-veined, 2-keeled,
keels with spinelike projections and often winged distally, smooth between
the keels, margins folded inward; upper glumes often shorter than
the lower glumes, 3-5-veined, keels entire; lower florets staminate; lower
paleas
present; upper florets bisexual or pistillate; lemmas and paleas
hyaline, unawned; anthers 3; style branches 2, red, free
to the base. Pedicels closely appressed but not fused to the rame
axes, flattened, thick, widening above the bases, glabrous. Pedicellate
spikelets usually
absent or rudimentary, occasionally well-developed. x = 9. Name
from the Greek eremos, solitary, and chloa, grass, a reference
to the solitary rame.
Eremochloa is a genus of 11 species that are native to Asia and Australia.
One species is naturalized in the southeastern United States; another was found
once in California but is not established in the Flora region.
1 |
Keels of the lower
glumes of the sessile spikelets winged distally, with 1-several
0.2-0.3 mm hooklike spines at the base ..... 1. E.
ophiuroides |
Keels of the lower glumes of the
sessile spikelets not winged, spine-bearing throughout, the
basal spines 1-3 mm ..... 2. E. ciliaris |
1. Eremochloa ophiuroides (Munro) Hack.
Centipede Grass
Plants mat-forming, stoloniferous, stolons to 150 cm, often branched, with
well-developed leaves, and (usually) axillary fascicles of closely imbricate leaves.
Culms 10-35 cm, unbranched. Sheaths mostly glabrous, margins sometimes
pilose, keeled; leaves mostly basal, blades of basal leaves 0.5-15
cm long, 1-5 mm wide, glabrous or pilose, with papillose-based hairs near the
base, margins glabrous or pectinate near the base; blades of upper leaves
reduced to obsolete. Rames 1-3, 3-12 cm, straight; internodes 2-2.3
mm. Sessile spikelets (2.2)3-4 mm long, (1.1)1.8-2.2 mm wide, elliptic;
calluses sparsely pubescent; glumes glabrous; lower glumes
5-7-veined, obtuse to truncate, often notched, keels with 1-several, 0.2-0.3 mm
hooklike spines near the base, winged distally; upper glumes 3-veined,
elliptic, acute; anthers of lower florets about 0.3 mm; anthers of upper
florets 1.5-1.7 mm. Pedicels 2.8-3.5 mm long, about 0.5 mm wide at
midlength. Pedicellate spikelets absent or to 3.4 mm, occasionally well-developed.
Caryopses 1.5-2 mm, purple to reddish-brown or brown. 2n = 18.
Eremochloa ophiuroides, an east Asian species, was introduced into the
southeastern United States as a lawn grass about 1920. It is now established along
roadsides and in woods, fallow fields, and dunes in the region. It flowers from
spring to fall, and sporadically at other times. The common name refers to the
appearance of the leafy stolons.
2. Eremochloa ciliaris (L.) Merr.
Plants cespitose, sometimes shortly stoloniferous. Culms 30-70
cm, compressed, sometimes branching from the upper nodes. Leaves mostly
basal;
sheaths keeled, strongly distichous and imbricate, often loose,
basal sheaths pubescent below; ligules to 0.6 mm, membranous; blades to
25 cm long, to 6 mm wide, flat or folded, basal blades glabrous. Rames 4-7
cm, straight to falcate; internodes 2-3 mm, clavate, shortly pubescent. Sessile
spikelets
4-5 mm, ovoid-oblong; calluses pubescent; lower glumes elliptic,
obscurely 7-9-veined, keeled, keels with conspicuous spines, basal spines
to 3 mm, those near the apices to 0.3 mm; upper glumes 3-5-veined,
keeled below;
lower florets staminate; anthers about 2 mm, yellow; upper
florets
bisexual. Pedicels to 3 mm. Pedicellate spikelets not differentiated.
2n = 36.
Eremochloa ciliaris is native to southeast Asia. It was collected in San
Francisco in the nineteenth century, but has not been reported since from the
Flora region.