| Mary E. Barkworth |
Plants annual or perennial; cespitose,
sometimes with extensive creeping stolons. Culms 15-200 cm. Leaves
usually not aromatic; ligules membranous, sometimes ciliate; blades
2-4 mm wide. Inflorescences terminal, sometimes also axillary but the
axillary inflorescences not numerous; peduncles with 1-many rames in
digitate or subdigitate clusters; rames sometimes naked basally, axes
terete to slightly flattened, without a translucent, longitudinal groove, bearing
1-many sessile-pedicellate spikelet pairs and a terminal triplet of 1 sessile
and 2 pedicellate spikelets, basal pair(s) homomorphic and homogamous, staminate
or sterile, unawned, persistent, distal spikelet pairs homomorphic but heterogamous,
sessile spikelets bisexual and awned, pedicellate spikelets staminate or sterile
and unawned; disarticulation in the rames, beneath the bisexual sessile
spikelets. Sessile spikelets often imbricate, dorsally compressed, with
blunt calluses; lower glumes chartaceous to cartilaginous, broadly convex
to slightly concave, sometimes pitted; lower florets reduced, sterile;
upper florets sterile or staminate and unawned in the homogamous pairs,
bisexual and awned in the heterogamous pairs; awns 1-3.5 cm, usually
glabrous; anthers (2)3. Pedicels free of the rame axes, terete
to somewhat flattened, slender, not grooved. Pedicellate spikelets sterile
or staminate. x = 10. Name from the Greek dicha, in two, as in
two separate things, and anthos, flower, a reference to the presence
of homogamous and heterogamous spikelets.
Dichanthium, a genus of 20 species, grows in habitats ranging from subdeserts
to marshlands in tropical Asia and Australia. It is frequently found in disturbed
areas, and some species are considered to provide good forage. Three species have
been introduced to the Flora region, one of which is sometimes used as
a lawn grass.
1 |
Lower glume of the sessile
spikelets with a subapical arch of hairs; pedicellate spikelets usually
sterile ..... 1. D. sericeum |
Lower glume of the sessile spikelets
without a subapical arch of hairs; pedicellate spikelets usually
staminate (2) |
|
Rame bases pilose; lower
glume of the sessile spikelets more or less obovate ..... 2. D.
aristatum |
|
Rame bases glabrous; lower glume of the
sessile spikelets elliptic to oblong ..... 3. D.
annulatum |
1. Dichanthium sericeum (R. Br.)
A. Camus
Queensland Bluegrass
Plants annual or perennial; tufted or cespitose. Culms 50-120 cm;
nodes densely pilose, hairs about 2 mm. Sheaths with scattered papillose-based
hairs; ligules 1-2 mm; blades 5-25 cm long, 2-5 mm wide. Rames
1-7, 3-7 cm, subdigitate, often glaucous and white-villous, spikelet-bearing to
the base, basal spikelet pairs consisting only of glumes; internodes pubescent,
hairs immediately below the nodes to 1.5 mm. Sessile bisexual spikelets
2.5-4.5 mm long, 1-1.4 mm wide; lower glumes 5-10-veined, with 0.7-1.5
mm hairs on the basal 1/2 and about 3 mm papillose-based hairs on the distal portion
of the keels and in a transverse subapical arch; awns 2-3.5 cm, twice-geniculate.
Pedicellate spikelets about 3 mm, usually sterile. 2n = 20.
Dichanthium sericeum is an Australian species. There are two subspecies:
D. sericeum (R. Br.) A. Camus subsp. sericeum is a perennial with
sessile spikelets 4-4.5 mm long and to 1-1.4 mm wide, 9-10-veined lower glumes,
and rames more than 4 cm long; D. sericeum subsp. humilius (J.M.
Black) B.K. Simon is an annual, with sessile spikelets up to 4 mm long and about
1 mm wide, 5-7-veined lower glumes, and rames less than 4 cm long. Dichanthium
sericeum subsp. sericeum is established in Texas and Florida.
2. Dichanthium aristatum (Poir.) C.E. Hubb.
Awned Dichanthium
Plants perennial; stoloniferous, stolons often 2 m or longer. Culms
70-100 cm, decumbent, erect portions generally about 35 cm, pubescent beneath
the inflorescences; nodes glabrous or densely short pubescent. Sheaths
glabrous; ligules 1-1.3 mm; blades 6-25 cm long, 3-6 mm wide,
glabrous or hispid. Rames (2)3-5(8), 4-7 cm, subdigitate, erect
to divergent, bases pilose, without spikelets; internodes pilose. Sessile
spikelets
4-5 mm; lower glumes more or less obovate, often involute, margins
ciliate basally, keels winged distally, apices obtuse; awns 1.5-2.5
cm, twice-geniculate.
Pedicellate spikelets 4-5 mm, usually staminate. 2n = 20.
Dichanthium aristatum was introduced to the Americas from southern Asia.
It is sometimes used as a lawn grass in Texas, Louisiana, and Florida.
3. Dichanthium annulatum (Forssk.) Stapf
Ringed Dichanthium
Plants perennial; stoloniferous. Culms to 100 cm, decumbent,
erect portions generally to 60 cm, often branched above the bases, glabrous
beneath the inflorescences; nodes glabrous or short-pubescent. Sheaths glabrous;
ligules 1-1.8 mm, truncate; blades 3-30 cm long, 2-7 mm wide,
scabrous, sparsely pilose, hairs sometimes papillose-based. Rames 2-9,
2.5-7 cm, subdigitate, erect to ascending, bases without spikelets, glabrous,
internodes ciliate on the margins. Sessile spikelets 2.5-5 mm long,
1-1.5 mm wide;
lower glumes elliptic or oblong, sparsely pubescent below, apices
obtuse, irregularly 2-3-toothed, 5-9-veined; upper glumes 3-veined; awns
1.3-2.2 cm, twice-geniculate. Pedicellate spikelets 2.5-5 mm, usually
staminate. 2n = 20, 40.
Dichanthium annulatum is native to southeastern Asia and is a highly esteemed
forage grass, especially in India. It is now established at scattered locations
in Texas and Louisiana.
2009, February: The records for Florida have been deleted because Bruce Hanson and Richard Wunderlin, after diligent searching were unable to find any specimens documenting the presence of the species in the state.