| Mary E. Barkworth |
Plants perennial, rarely annual;
cespitose, loosely tufted, or mat-forming, sometimes rhizomatous or stoloniferous.
Culms 7-300 cm, not woody, often decumbent at the base, erect to ascending.
Sheaths open; ligules membranous, truncate, ciliate; blades
flat or convolute, usually obtuse. Inflorescences terminal, sometimes
also axillary, panicles of 2-many, digitately, subdigitately, or racemosely
arranged spikelike branches; branches triquetrous, spikelets subsessile
or sessile, solitary, in 2 rows, lower lemmas appressed to the branch axes;
disarticulation below the glumes. Spikelets dorsally compressed,
with 2 florets; lower florets sterile or staminate; upper florets
sessile, bisexual. Lower glumes absent; upper glumes and lower
lemmas equal, membranous; lower paleas absent; upper lemmas
indurate, usually glabrous, sometimes with an apical tuft of hairs, margins
slightly involute, clasping the palea, apices acute to obtuse; upper paleas
similar to the upper lemmas in texture. Caryopses ellipsoid. x
= 10. Name from the Greek axon, axis, and pes, foot.
Axonopus is a genus of approximately 100 tropical and subtropical species,
most of which are native to the Western Hemisphere. Three species are native
to the Flora region; one additional species has been grown experimentally
in Florida.
All the species tend to grow in open habitats, often where the soil is somewhat
impermeable and slightly flooded in the rainy season. Axonopus
fissifolius and A. compressus are cultivated for forage in many countries; A. compressus is also
used as a lawn grass. Both species are inclined to be weedy. The presence of
rhizomes or stolons is affected by environmental conditions, with plants growing
in crowded conditions, e.g., lawns, rarely producing them.
1 |
Panicles
with 30-100+ branches; lower branches 10-24 cm long; culms (50)100-300
cm tall ..... 4. A. scoparius |
Panicles with 2-7 branches; lower branches
1-15 cm long; culms 7-100 cm tall (2) |
|
Spikelets 3.5-5.5 mm
long; upper glumes glabrous; lower lemmas glabrous or sparsely pilose
over the veins ..... 3. A. furcatus |
|
Spikelets 1.6-3.5 mm long; upper
glumes and lower lemmas sparsely pilose on the margins or marginal veins (3) |
|
Upper glumes and lower
lemmas extending beyond the upper florets, forming acute to acuminate
apices; blades 3-20 mm wide ..... 2. A.
compressus |
|
Upper glumes and lower lemmas not or
scarcely extending beyond the upper florets, forming obtuse to subacute
apices; blades 1.5-6 mm wide ..... 1. A.
fissifolius |
1. Axonopus fissifolius (Raddi) Kuhlm.
Common Carpetgrass
Plants usually cespitose, sometimes stoloniferous, nodes of the stolons
often pilose. Culms 10-75 cm, erect or depressed-decumbent; cauline
nodes glabrous or slightly pubescent. Sheaths compressed, mostly glabrous,
margins ciliate; ligules 0.2-0.4 mm; blades 4-15 cm long, 1.5-6
mm wide, flat, mostly glabrous, margins with papillose-based cilia. Panicles
terminal and axillary, 5-11 cm overall, rachises to 3 cm, with 2-7 branches; branches
2-9(12) cm, spreading or ascending. Spikelets 1.6-2.2(2.8) mm, ovoid or
ellipsoid, obtuse to acute. Upper glumes and lower lemmas scarcely
extending beyond the upper florets, 2-veined, margins sparsely pilose, apices
obtuse to subacute; upper lemmas and paleas 1.6-2.1 mm long, 0.5-0.7
mm wide. Caryopses 1.5-1.8 mm, gray. 2n = 20, 40, 80, 100.
Axonopus fissifolius is sometimes used as a lawn or pasture grass, but
it is also an invasive weedy species, often growing in moist, disturbed sites.
It is native in the southeastern United States and from central Mexico south to
Bolivia and Argentina. It has also been introduced into tropical and subtropical
regions of the Eastern Hemisphere.
2. Axonopus compressus (Sw.) P. Beauv.
Broadleaf Carpetgrass
Plants stoloniferous, rarely rhizomatous, rhizomes, when present, 3-5 cm.
Culms 7-80 cm; nodes glabrous or pubescent. Sheaths keeled,
strongly compressed, pubescent; ligules 0.3-0.5 mm; blades 3-20
mm wide, glabrous or sparsely pilose, midveins often white and prominent, apices
frequently ciliate or pubescent. Panicles terminal and axillary, 4-10 cm
overall, rachises to 3.5 cm, with 2-5 branches; branches 1-13 cm. Spikelets
2-3.5 mm, ovoid, ellipsoid, or lanceoloid, acuminate. Upper glumes and
lower lemmas extending beyond the upper florets, 2-5-veined, marginal veins
pilose, apices acute to acuminate; upper lemmas and paleas 1.5-1.8
mm, broadly ellipsoid. Caryopses 1.2-1.5 mm, gray. 2n = 40, 60,
80.
Axonopus compressus is native from the southeastern United States to Bolivia,
Brazil, and Uruguay, and has become established in the Eastern Hemisphere. It
is used as a lawn and forage grass but is also weedy, readily growing in moist,
disturbed habitats.
3. Axonopus furcatus (Flüggé)
Hitchc.
Big Carpetgrass
Plants stoloniferous. Culms 30-100 cm; nodes glabrous or pubescent.
Sheaths compressed, glabrous or sparsely to densely pilose, hairs appressed;
ligules 0.3-1 mm; blades 3-25 cm long, 2-15 mm wide, margins often
with papillose-based hairs near the base, scabrous distally. Panicles terminal
and axillary, with 2(-4) divergent branches; branches 4-15 cm. Spikelets
3.5-5.5 mm long, about 1.5 mm wide, sessile or subsessile, ovoid-ellipsoid, acuminate.
Upper glumes glabrous, 5-7-veined; lower lemmas 5-7-veined, glabrous
or sparsely pilose over the veins; upper lemmas and paleas 2.5-3.2
mm, light yellow, obtuse. Caryopses 1.8-2.2 mm, obovate, yellow. 2n
= unknown.
Axonopus furcatus is endemic to the southeastern United States. It grows
in moist pine barrens, marshes, river banks, wet ditches, pond margins, and other
such damp areas.
4. Axonopus scoparius (Humb. & Bonpl. ex
Flüggé) Kuhlm.
Carpetgrass
Plants rhizomatous, frequently also stoloniferous. Culms
(50)100-300 cm, to about 1 cm thick, sometimes branching above the base; nodes
glabrous. Leaves primarily cauline; sheaths often much wider than
the internodes, mostly glabrous but the collars pubescent, lower sheaths compressed;
ligules 0.5-2.9 mm, ciliolate; blades 15-50 cm long, (5)20-35
mm wide, bases usually wider than the sheaths. Panicles terminal and
axillary, 10-50 cm overall, rachises 2-3.5 cm, with 30-100+ branches; lower
branches 10-24 cm, frequently fascicled. Spikelets 2.1-2.7 mm, ovoid
to oblong-ellipsoid, acute or apiculate; upper glumes and lower lemmas
usually 5-veined, sparsely pilose; upper florets 0-0.4 mm shorter than
the upper glumes and lower lemmas, obtuse to subacute. Caryopses usually
absent. 2n = 20.
Axonopus scoparius is native from southern Mexico to Peru, Bolivia, and
Brazil. In Mesoamerica, it rarely sets seed but is grown for forage and often
persists after cultivation has ceased. It has been grown experimentally in Florida,
but it is not winter hardy even there. Not surprisingly, A. scoparius is
not established in the Flora region.