| Nassella:
Species diversity and distribution Barkworth, M.E. and M.A. Torres. 2001. Taxon 50:439-468. |
| Summary; Introduction; Materials and Methods; Results; Species list; Excluded species; Distribution by country; Distribution by species; Distribution changes; Excluded species; Discussion. |
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Species Listing The rectangular box that appears in some entries stands for the identity symbol; names following it are nomenclatural synonyms of the accepted name. Nassella airoides (E. Ekman) Barkworth [≡ Stipa airoides Ekman]. Isotypes: BAA ex S, Col. Typus 2944! CORD! Nassella ancoraimensis F. Rojas Nassella arcaensis (Speg.) Torres [≡ Stipa arcaensis Speg.]. Lectotype: LP ex LPS 2457! [= Stipa paniculate Hitchc., Isotypes: LIL! LP!]. Nassella arcuata (R. E. Fries) Torres [≡ Stipa arcuata R. E. Fries]. Isotype: Fries 944 CORD! Nassella arechavaletae (Speg.) Barkworth [≡ Stipa arechavaletae Speg.]. Syntype: LP ex LPS 2456!, Isosyntype: BAA! Nassella argentinensis (Speg.) Peñailillo [≡ Stipa argentinensis Speg.]. Holotype: LP ex LPS 2458! [= Stipa neesiana var. virescens Hack., Isotype: BAA Col. Typus 3181!]. Nassella asplundii Hitchc. Holotype: US 1099355! Nassella ayacuchensis (Tovar) Barkworth [≡ Stipa ayacuchensis Tovar]. Holotype: US 3029243! Nassella brachychaetoides (Speg.) Barkworth [≡ Stipa brachychaetoides Speg.]. Holotype: LP ex LPS 2459!, Isotype: BAA! Nassella brachyphylla (Hitchc.) Barkworth [≡ Stipa brachyphylla Hitchc.]. Holotype: US 1099323! Nassella brasiliensis (A. Zanín & Longhi-Wagner) Peñailillo [≡ Stipa brasiliensis A. Zanín & Longhi-Wagner]. Nassella burkartii (Torres) Barkworth & Torres. Nassella cabrerae Torres. Holotype: Cabrera 9299 LP!, Isotype BAA! Nassella caespitosa Griseb. [≡ Stipa caespitosa (Griseb.) Speg.]. Isotypes: Lorentz 78 BAA!, CORD! [ = Stipa caespitosa var. subtypica Speg. (1901), nom. inval.; = Stipa plagiostephana Speg.; = Stipa violacea Hitchc.]. Nassella carettei (Hauman) Torres [≡ Stipa carettei Hauman]. Syntype: BAA Col. Typus 2998! Nassella catamarcensis Torres. Holotype: Parodi 14174 BAA! Nassella cernua (Stebbins & Love) Barkworth [≡ Stipa cernua Stebbins & Love; ≡ Stipa pulchra var. cernua (Stebbins & Love) Beetle & Tofsrud]. Holotype: UC 641631! Nassella chaparensis F. Rojas Holotype: LPB. Nassella charruana (Arechav.) Barkworth [≡ Stipa charruana Arechav.; = Stipa longicylindrica Kuntze]. Note: Introduced to Australia (Jacobs & al. 1998). Nassella chilensis (Trin.) E. Desv. [≡Urachne chilensis Trin.; ≡ Piptatherum ramosum Nees p.p. incl. leg. Bertero; ≡ Nassella ramosa (Nees) Desv.; ≡ Urachne ramosa (Nees) Steud.; nom. inval.; Urochloa gigantea Steud.; = Stipa landbeckii var. eremophila Phil.; = Piptatherum lindleyanum Nees; = Nassella laxiflora Phil.; = Nassella sadae Phil.; = Nassella planifolia Phil.; = Nassella floribunda Phil.; = Nassella obscura F. Phil.]. Nassella clarazii (Ball) Barkworth [≡ Stipa clarazii Ball]. Type fragment: BAA Col.Typus 3003 (p.p.a.)! [=Stipa quadrifaria Kuntze; =Stipa clarazii var. bulbosa Speg.]. Nassella
coquimbensis (Matthei) Peñailillo
[≡ Stipa coquimbensis Matthei]. Isotype: CONC 29361! This
species is unusual in Nassella in having long hairs on the awn
column, a 2-veined palea 1/3-1/2 as long as the anthoecium, and short
rhizomes, but we agree with Peñailillo that it belongs in Nassella. Nassella cordobensis (Speg.) Barkworth [≡ Stipa cordobensis Speg.]. Lectotype: LP ex LPS 2451!; Syntype: LP ex LPS 2452! [= Stipa tenuis var. papillosa Hack.; = Stipa papillosa (Hack.) Hitchc.; = Stipa cordobensis var. tuberculata F. A. Roig]. Nassella crassiflora (Roseng. & B. R. Arill.) Barkworth [≡ Stipa crassiflora Roseng. & B. R. Arill.]. Isotype: BAA Col. Typus 3010! Nassella curamalalensis (Speg.) Barkworth [≡ Stipa curamalalensis Speg.; ≡ Stipa tenuissima var. curamalalensis (Speg.) Hack.]. Holotype: LP ex LPS 2455! Nassella curviseta (Hitchc.) Barkworth [≡ Stipa curviseta Hitchc.]. Type: US 1099338! Nassella dasycarpa (Hitchc.) Torres [≡ Stipa dasycarpa Hitchc.]. Holotype: US 1099337! Nassella
depauperata (Pilg.) Barkworth [≡ Stipa
depauperata Pilg.]. Isotype: Weberbauer
6903 BAA! [= Stipa
leptothera var. atroviolacea
Hack., Type: W!; = Stipa
depauperata var. humilis Pilger]. Nassella duriuscula (Phil.) Barkworth [≡ Stipa duriuscula Phil.]. Type fragment: BAA Col. Typus 3037 ex SGO! Nassella elata (Speg.) Torres [≡ Stipa caespitosa var. elata Speg.]. Holotype: LP ex LPS 12524! Nassella entrerriensis (Burkart) Peñailillo [≡ Stipa entrerriensis Burkart]. Holotype: Jozami 8 SI! Nassella fabrisii Torres. Holotype: Fabris 1374 LP! Nassella famatinensis Torres. Holotype: J. Hunziker 1965 BAA! Nassella filiculmis (Delile) Barkworth [≡ Stipa filiculmis Delile; = Stipa leucogluma Steud.; = Stipa ceresiensis Kuntze; Stipa filiculmis var. major Speg.; = Stipafiliculmis var. minor Speg.,]. Note: Introduced to the south of France, probably in wool (Thellung 1912). Nassella formicarum (Delile) Barkworth [≡ Stipa formicarum Delile]. Type fragment: BAA Col. Typus 3055 ex Herb. Montpellier! Note: Introduced to the south of France, probably in wool (Thellung 1912). Nassella gibba (Phil.) M. Muñoz [≡ Piptochaetium gibbum Phil.; = Nassella exserta var. asperata Parodi fide Muñoz-Schick (1990)]. Nassella gigantea (Steud.) M. Muñoz [≡ Urachne gigantea Steud.; = Nassella exserta Phil.; = Nassella melanocarpa Phil. = Nassella fuscescens Phil.; = Nassella pugae Phil. fide Muñoz-Schick (1990)]. Nassella glabripoda Torres. Holotype: LIL 14901!; Isotype: BAA! Nassella hirtifolia (Hitchc.) Barkworth [≡ Stipa hirtifolia Hitchc.]. Nassella holwayii (Hitchc.) Barkworth [≡ Stipa holwayii Hitchc.]. Paratype fragment: BAA Col. Typus 4166! Nassella huallancaensis (Tovar) Barkworth [≡ Stipa huallancaensis Tovar]. Nassella hunzikeri (Caro) Barkworth [≡ Stipa hunzikeri Caro]. Holotype: A. T. Hunziker 7191 CORD! Nassella hyalina (Nees) Barkworth [≡ Stipa hyalina Nees]. Type fragment: BAA Col. Typus 3088 ex B! Note: Introduced to New South Wales, Australia (Vickery & al. 1986; Jacobs & al. 1998). Nassella ibarrensis (Kunth) Laegaard, non Peñailillo, nom. illeg. [≡ Stipa ibarrensis Kunth]. Holotype: P! Note: Peñailillo published this combination in Gayana, Bot. 55(2): 86. The title page gives a date of 1998 but this volume was not actually published until May, 1999, so Laegaard’s combination, which really was published in 1998, has priority. Nassella inconspicua (J. Presl) Barkworth [≡ Stipa inconspicua J. Presl]. Type fragment: BAA Col. Typus 3099 ex PR! [= Urachne haenkeana Trin & Rupr., nom inval.; = Stipa sodiroana Hack. fide Tovar (1993); = Stipa caespitosa var. lilloi Hack., Isotype: LIL 39986!]. Nassella juergensii (Hack.) Barkworth [≡ Stipa juergensii Hack.]. Type fragment: BAA Col. Typus 3108! Nassella juncea Phil. [≡ Nassella chilensis var. juncea (Phil.) M. Muñoz]. Nassella karstenii (Hitchc.) Peñailillo [≡ Stipa karstenii Hitchc.]. Holotype: US 1126745! Nassella lachnophylla (Trin.) Barkworth [≡ Stipa lachnophylla Trin.; ≡ Stipa cumingiana var. lachnophylla (Trin.) Trin. & Rupr.; Stipa cumingiana Trin.]. Nassella laevissima (Phil.) Barkworth [≡ Piptochaetium laevissimum Phil., ≡ Stipa laevissima (Phil.) Speg., = Stipa breviculmis Hitchc., = Stipa lahittei Parodi]. Nassella lepida (Hitchc.) Barkworth [≡ Stipa lepida Hitchc.]. Holotype: US 733683! [= Stipa eminens var. andersonii Vasey, Holotype US 556912!; = Stipa lepida var. andersonii (Vasey) Hitchc.; Stipa hassei Vasey]. Nassella leptocoronata (Roseng. & B. R. Arill.) Barkworth [≡ Stipa leptocoronata Roseng. & B. R. Arill.]. Isotype: BAA ex MVFA 2145! Nassella leptothera (Speg.) Torres [≡ Stipa leptothera Speg.]. Lectotype: LP ex LPS 12642! Nassella leucotricha (Trin. & Rupr.) R. W. Pohl [≡ Stipa leucotricha Trin. & Rupr.]. Note: Recorded as being introduced into Australia (Vickery et al. 1986). Nassella ligularis (Griseb.) Barkworth & Torres. [= Stipa neesiana var. ligularis Griseb, S. ligularis (Griseb.) Speg.]. Isotype CORD! Nassella linearifolia (E. Fourn.) R. W. Pohl [≡ Stipa linearifolia E. Fourn.; = Oryzopsis florulenta Pilg.; = Stipa florulenta (Pilg.) Parodi]. Nassella longicoronata (Roseng. & B. R. Arill.) Barkworth [≡ Stipa longicoronata Roseng. & B. R. Arill.]. Holotype fragment: SI ex MVFA 2150! Nassella longiglumis (Phil.) Barkworth [≡ Stipa longiglumis Phil.]. Probable type fragment: SGO 45631! Nassella macrathera (Phil.) Barkworth [≡ Stipa macrathera Phil.]. Probable isotype fragment: BAA Col. Typus 3130 ex SGO! Nassella manicata (E. Desv.) Barkworth [≡ Stipa manicata E. Desv.]. Type fragment: BAA Col. Typus 3135 ex P! Note: Introduced and established in California in areas associated with sheep grazing; misidentified as N. formicarum by Barkworth (1993b). Nassella megapotamica (Spreng. ex Trin.) Barkworth [≡ Stipa megapotamica Spreng. ex Trin.; = Stipa latifolia Hack. & Arechav.; = Stipa latissimifolia Kuntze; = Stipa manicata var. latifolia (Hack. & Arechav.) Speg.; = Stipa manicata var. media Speg.]. Note: Established, but apparently not spreading, near Canberra, Australia (Jacobs & Everett 1993; Jacobs & al. 1998). Nassella melanosperma (J. Presl) Barkworth [≡ Stipa melanosperma J. Presl; = Stipa bonariensis Henrard & Parodi; = Nassella bonariensis (Henrard & Parodi) Barkworth]. Nassella mexicana (Hitchc.) R. W. Pohl [≡ Stipa mexicana Hitchc.]. Holotype: US 993386! Nassella meyeniana (Trin. & Rupr.) Parodi [≡ Urachne meyeniana Trin. & Rupr.; = Urachne laevis Trin. & Rupr.; = Piptatherum ramosum Nees p.p. excl. leg. Bertero; = Nassella corniculata Hack. ]. Nassella meyeri Torres. Holotype: Meyer 14886 LIL!; Isotype: BAA! Nassella mucronata (Kunth) R. W. Pohl [≡ Stipa mucronata Kunth]. Lectotype fragment: US 294533 ex P! [= Stipa grisebachii E. Fourn.; = Stipa setigera J. Presl in C. Presl (1830); = Stipa trochlearis Nees & Meyen]. Note: Introduced to Coimbra, Portugal and Aranjuez, Spain (Vázquez & Devesa 1996), France (Kerguélen 1975; Martinovsky 1980), and Italy (Martinovsky 1980; Pignatti 1982; Moraldo 1986). Nassella nardoides (Phil.) Barkworth [≡ Danthonia nardoides Phil.; ≡ Stipa nardoides (Phil.) Hack. ex Hitchc.]. Isotype fragment: BAA Col. Typus 3176! [= Stipa calchaquia Speg.]. Nassella neesiana (Trin. & Rupr.) Barkworth [≡ Stipa neesiana Trin. & Rupr.; = Stipa eminens Nees, non Cav.; = Stipa neesiana var. chilensis Trin. & Rupr.; = Stipa neesiana var. fernandeziana Trin. & Rupr.; = Stipa skottsbergii Pilger; = Stipa longiflora Steud.; = Stipa trachysperma Phil.; = Stipa fernandeziana Phil.; = Stipa hispida Phil., = Stipa neesiana var. glabrata Arechav.; = Stipa neesiana var. hirsuta Arechav.; = Stipa neesiana var. longiaristata Arechav.; = Stipa sublaevis Speg.; = Stipa setigera var. glabrata Arechav. ex Speg.; = Stipa setigera var. hispidula Speg.]. Note: Introduced into southern France (Thellung 1912), New South Wales and Victoria, Australia (Vickery & al. 1986; Jacobs & Everett 1993, 1996); and New Zealand (Jacobs & al. 1989; Connor & al. 1993; Edgar & Connor 2000). Nassella nidulans (Mez) Barkworth [≡ Stipa nidulans Mez]. Isotype: Hieronymus 298 CORD! Nassella niduloides (Caro) Barkworth [≡ Stipa niduloides Caro]. Holotype: A. T. Hunziker 9553 CORD! Nassella novarae Torres. Holotype: Cabrera 18524 LP! Nassella nubicola (Speg.) Torres [≡ Stipa nubicola Speg.]. Lectotype: LP ex LPS 2426! Nassella nutans (Hack.) Barkworth [≡ Stipa nutans Hack.]. Holotype fragment: BAA Col. Typus 3184 ex US! Nassella pampagrandensis (Speg.) Barkworth [≡ Stipa pampagrandensis Speg.]. Lectotype: LP ex LPS 2419! Nassella pampeana (Speg.) Barkworth [≡ Stipa pampeana Speg.]. Holotype: LP ex LPS 2420; Isotype: BAA! Nassella paramilloensis (Speg.) Torres [≡ Stipa paramilloensis Speg.]. Lectotype: LP ex LPS 2422! [= Stipa perrigida Speg., Holotype: LP ex LPS 2423!; = Stipa gilliesii Hitchc., Isotype BAA Col. Typus 3061!]. Nassella parodii (Matthei) Barkworth [≡ Stipa parodii Matthei]. Nassella parva Torres. Holotype: LIL 513135! Nassella pauciciliata (Roseng. & Izag.) Barkworth [≡ Stipa megapotamica var. pauciciliata Roseng. & Izag.]. Isotype: BAA Col. Typus 3162! [≡ Stipa pauciciliata (Roseng. & Izag.) Roseng., Arill., & Izag.]. Nassella pfisteri (Matthei) Barkworth [≡ Stipa pfisteri Matthei]. Nassella philippii (Steud.) Barkworth [≡ Stipa philippii Steud.]. Type fragment: BAA Col. Typus 3214! [= Stipa bavioensis Speg.; = Stipa bavioensis var. minor Speg., Holotype: LP ex LPS 2463!]. ]. Nassella
pittieri (Hitchc.) Peñailillo
[≡ Stipa pittieri Hitchc.]. Holotype: US 531634! Nassella planaltina (A. Zanín & Longhi-Wagner) Peñailillo [≡ Stipa planaltina A. Zanín & Longhi-Wagner]. Paratype: UTC 219007! Nassella poeppigiana (Trin. & Rupr.) Barkworth [≡ Stipa poeppigiana Trin. & Rupr.; = Stipa amethystina Steud.; = Stipa manicata var. typica Speg., nom. inval.; = Stipa latifolia var. grandiflora Hack. in Stuck.; = Stipa megapotamica var. latiflora Roseng. & Izag.; = Nassella amethystina (Steud.) Barkworth]. Nassella pseudopampagrandensis (Caro) Barkworth [≡ Stipa pseudopampagrandensis Caro]. Syntype: CORD ex Herb. Stuckert 16804! Nassella psittacorum (Speg.) Peñailillo [≡ Stipa psittacorum Speg.]. Holotype: LP ex LPS 2445! Nassella pubiflora (Trin. & Rupr.) E. Desv. [≡ Urachne pubiflora Trin. & Rupr., ≡ Oryzopsis pubiflora (Trin. & Rupr.) Scribn.; ≡ Stipa pubiflora (Trin. & Rupr.) M. Muñoz.; = Nassella flaccidula Hack.; = Nassella deltoidea Hack.]. Nassella pulchra (Hitchc.) Barkworth [≡ Stipa pulchra Hitchc.]. Holotype: US 416590! Nassella punensis Torres. Holotype: Cabrera & al. 19778 LP!; Isotype: BAA! Nassella pungens E. Desv. [= Nassella humilis Phil.; = Nassella johnstonii Parodi]. Nassella ragonesei Torres. Holotype: Ragonese & Piccinini 9777 BAA! Nassella rhizomata (A. Zanín & Longhi-Wagner) Peñailillo [≡ Stipa rhizomata A. Zanín & Longhi-Wagner]. Paratype: UTC 219006! Nassella rosengurttii (Chase) Barkworth [≡ Stipa rosengurttii Chase]. Isotype: BAA Col. Typus 3234! Nassella rupestris (Phil.) Torres [≡ Stipa rupestris Phil.]. Type: SGO 63159! Nassella sanluisensis (Speg.) Barkworth [≡ Stipa sanluisensis Speg.]. Lectotype: LP ex LPS 12646! [= Stipa debilis Mez; = Stipa debilis var. papillosa Caro; = Stipa sanluisensis var. longispicula Caro]. Nassella sellowiana (Nees ex Trin. & Rupr.) Peñailillo [≡ Stipa sellowiana Nees ex Trin. & Rupr.]. Type fragment: BAA Col. Typus 3254 ex B! [= Stipa molfinoi Speg. Holotype: LP ex LPS 2418!]. Nassella smithii (Hitchc.) Barkworth [≡ Stipa smithii Hitchc.]. Holotype: US 1357804! Nassella soukupii (Tovar) Barkworth [≡ Stipa soukupii Tovar]. Isoype: Tovar 2814 LP! Nassella spegazzinii (Arechav.) Barkworth [≡ Stipa spegazzinii Arechav.; = Stipa uruguaycola Speg. Lectotype: LP ex LPS 2427!]. Nassella stuckertii (Hack.) Barkworth [≡ Stipa stuckertii Hack.]. Isotype: CORD! Nassella
subnitida (Roseng. & B. R. Arill.) Barkworth [≡ Stipa
subnitida Roseng. & B. R. Arill.]. Isotype: BAA Col. Typus 3276! Nassella tenuiculmis (Hack.) Peñailillo [≡ Stipa tenuiculmis Hack.]. Nassella tenuis (Phil.) Barkworth [≡ Stipa tenuis Phil.]. Type fragment: LP ex LPS 22996! [= Stipa argentina Speg.; = Stipa tenuis var. argentina (Speg.) Speg.; = Stipa puelches Speg.]. Nassella tenuissima (Trin.) Barkworth [≡ Stipa tenuissima Trin.; = Stipa cirrosa E. Fourn.; = Stipasubulata E. Fourn.; = Stipa oreophila Speg.; = Stipa tenuissima var. oreophila (Speg.) Speg.; = Stipa tenuissima var. planicola Speg.]. Note: Introduced to Australia (Jacobs & al. 1998), New Zealand (Ford 1994; Connor & Ford 1996; Gardner & al. 1996), and South Africa (Connor & Ford 1996). Cultivated in California (Barkworth, pers. obs.) and New Zealand (Connor & Ford 1996; Edgar & Connor 2000). Nassella torquata (Speg.) Barkworth [≡ Stipa torquata Speg.]. Holotype: LP ex LPS 2429! Nassella trachyphylla Henrard Nassella trichotoma (Nees) Hack. ex Arechav. [≡ Stipa trichotoma Nees; ≡ Agrostis trichotoma (Nees) Nees ex Trin.; ≡ Urachne trichotoma (Nees) Trin.; ≡ Piptochaetium trichotomum (Nees) Griseb.; ≡ Oryzopsis trichotoma (Nees) Druce.]. Type fragment: BAA Col. Typus 3292 ex B! Note: Introduced to southern France and Italy (Burkart 1969), Australia and New Zealand (Vickery & Jacobs 1980; Jacobs & Everett 1993, 1996; Jacobs & al. 1989; Edgar & Connor 2000), and the U.S.A. (U.S.D.A.-A.P.H.I.S. 1990). So far as is known, all populations of the species have been eliminated from the continental U.S.A. The U.S. Department of Agriculture regards the species as one of the ten worst potential weeds for U.S. agriculture. Nassella
tucumana (Parodi) Torres [≡ Stipa
tucumana Parodi]. Holotype: Parodi 11062 BAA! [= Nassella asperifolia F. Rojas, Isotype: UTC 203411!]. Nassella uspallatensis (Speg.) Torres [≡ Stipa uspallatensis Speg.]. Syntype: LP ex LPS 12649! Nassella
vallsii
(A. Zanín & Longhi-Wagner) Peñailillo [≡ Stipa
vallsii A. Zanín & Longhi-Wagner]. Paratype: UTC 219008! Nassella vargasii (Tovar) Peñailillo [≡ Stipa vargasii Tovar]. Holotype: US 2474222! Nassella ventanicola (Cabrera & Torres) Barkworth [≡ Stipa ventanicola Cabrera & Torres]. Holotype: Cabrera 7305 LP! Nassella viridula (Trin.) Barkworth [≡ Stipa viridula Trin.]. Holotype: LE! [= Stipa parviflora Nutt., non Desf., = Stipa nuttalliana Steud.]. Nassella wurdackii (Tovar) Barkworth [≡ Stipa wurdackii Tovar]. Nassella yaviensis Torres. Holotype: Cabrera & al. 15357 LP! |