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Preparation of the following publications drew on the resources and/or facilities of the Intermountain Herbarium to some extent.  The list is incomplete, as I have not been tracking such publications sufficiently diligently of late.  The listing starts with 1996.  

Web Pages are listed separately because they are probably updated at irregular intervals. Some may have become extinct. 

Mary Barkworth, Director.

Print or CD Publications 

Articles in scientific journals and books are in black; other scientific publications (theses, reports, etc.) are in green; popular articles are in blue; abstracts in grey..

Years available: 1996 1997 1998 1999 2000 2001 2002 2003, 2004, 2005


Patterson, Jason T., Steven R. Larson, and Paul G. Johnson. 2005. Genome relationships in polyploid Poa pratensis and other Poa species inferred from phhylogenetic anaylsis of nuclear and chloroplast DNA. Genome 48:76-87.



Barkworth, Mary. E., Kathleen M. Capels, Sandy Long, Michael B. Piep. 2003. Flora of North America, volume 25. Poaceae, part 2. Oxford University Press, New York. 783 pp.

Gandhi, K. and M.E. Barkworth. 2003. Proposal to conserve the name Munroa (Gramineae) with that spelling. Taxon 52:??–??.


Barkworth, M.E. and S.W.L. Jacobs. 2002. Voucher specimens: a critical element in a scientific study.  Pp. 112-118 in P. Hernández, M.T. Moreno, J.I. Cubero, and A. Martin (Editors),  Proceedings of the Fourth International Triticeae Symposium. Junta de Andalucia, Consejería de Agricultura y Pesca, Córdoba, Spain.

Miller, C.F. and M. Kim.  2002.  Ilex x wandoensis C.F. Miller and M. Kim, a new hybrid species of Ilex (Aquifoliaceae) from Korea.  Korean Journal of Plant Taxonomy 32 (3):  293-299.

Leidolf, A., S. McDaniel, and T. Nuttle. 2002. The flora of Oktibbeha County, Mississippi. Sida 20:691-765.

Panter, K.E., Manners, G.D., Stegelmeier, B.L., Lee, S., Gardner, D.R., Ralphs, M.H., Pfister, J.A. and James, L.F.  Larkspur Poisoning:  Toxicology and alkaloid structure-activity relationships.  Biochemical Systematics and Ecology 30:113-128, 2002.

Panter, K.E., James, L.F., Gardner, D.R., Ralphs, M.H., Pfister, J.A., Stegelmeier, B.L. and Lee, S.T.  Reproductive losses to poisonous plants:  Influence on management strategies.  J. Range Manage. 55(3), 301-308, 2002.

Pfister, J.A., D.R. Gardner, B.L. Stegelmeier, A.P. Knight, J.W. Waggoner, Jr., and J.O. Hall.  Plains larkspur (Delphinium geyerii) grazing by cattle in Wyoming.  J. Range Manage.  55:  In press.

Valdes-Reyna, J. and M.E. Barkworth. 2002. Flora of Veracruz, fasc. 127: Poaceae II, Pooideae: Tribu Stipeae. 28 pp.


Archibald J.K., P.G. Wolf, V. J. Tepedino, and J. Bair. 2001. Genetic relationships and population structure of the endangered steamboat buckwheat, Eriogonum ovalifolium var. williamsiae (Polygonaceae). American Journal of Botany, 88: 608-615.

Barkworth, M.E. and S.W.L. Jacobs. 2001. Valuable research or short stories: what makes the difference? Hereditas 135:263-270.

Barkworth, M.E. and M.A. Torres. 2001. Distribution and diagnostic characters of Nassella (Poaceae: Stipeae). Taxon 50:439-468.

Benson, L., J. Daniel, W. Forsyth.  2001.  Plants of Cache Valley. [interactive multimedia CD].

Bosch, J. and William Kemp. 2001. How to manage the blue orchard bee as an orchard pollinator. Sustainable Agriculture Network, Beltsville, Maryland 20705-2351.

Flora ID Northwest. 2001.  Plants of Utah: Interactive keys and color photos, vers. 4.  New York Botanical Garden Press. [includes pictures, some from the Intermountain Herbarium Slide Collection].  

Ivie, M.A. 2001.  On the geographic origin of the wheat stem sawfly (Hymenoptera: Cephidae): a new hypothesis of introduction from northeastern Asia.  American Entomologist 47:84-97.

Kropp, B. R. 2001. Familiar faces in unfamiliar places: mycorrhizal fungi associated with Caribbean pine. Mycologist 15: 137-140.

Larson, S.R., B. Waldron, S. Monson, L. St. John, A.L. Palazzo, C. L. McCracken, and D. Harrison.  2001.  Patterns of AFLP variation in the apomictic Poa secunda Presl. complex of western North America.  Crop Science 41, In Press .

Massa A.N., S.R. Larson, K.B. Jensen, and D.J. Hole..  2001.  AFLP variation in Bromus section Ceratochloa germplasm of Patagonia.  Crop Science 41:1609-1616.

Matheny, P.B. and B.R. Kropp.  A revision of the Inocybe lanuginosa group and allied species in North America.  Sydowia 53(1):93-139.

Panter, K.E., Mayland, H.F., Gardner, D.R. and Shewmaker, G. 2001.  Beef cattle losses after grazing Lupinus argenteus (Slivery Lupine).  Vet. and Human Toxicol. 43 (5) :279-282.

Panter, K.E, G.D. Manners, B.L. Stegelmeier, S.T. Lee, D.R. Gardner, M.H. Ralphs, J.A. Pfister, and L.F. James.  2001.  Larkspur poisoning:  toxicology and alkaloid structure-activity relationships.  Biochem. System. Ecol.  In press.

Pierson, K., V. J. Tepedino, S. Sipes, and K. Kuta.  2001.  Pollination ecology of the rare orchid, Spiranthes diluvialis:  implications for conservation.  Proc. 3rd Southwest Rare and Endangered Plant Conf., Flagstaff, AZ.  In Press.

Pfister, J.A., M.H. Ralphs, D.R. Gardner, B.L. Stegelmeier, G.D. Manners, K.E. Panter, and S.T. Lee.  2001.  Management of three toxic Delphinium species based on alkaloid concentrations. Biochem. System. Ecol. In press.

Pfister, J.A., B.L. Stegelmeier, C.D. Cheney, M.H. Ralphs, and D.R. Gardner.  2001.  Conditioning taste aversions to locoweed (Oxytropis sericea) in horses.  J. Anim. Sci. 79: In press.

Pryer, K. M., H. Schneider, A. R. Smith, R. Cranfill, P. G. Wolf, J. S. Hunt, and S. D. Sipes. 2001. Horsetails and ferns are a monophyletic group and the closest relatives to seed plants. Nature 409: 618-622

Smith, Alan R., H. Tuomisto, K.M. Pryer, J.S. Hunt, and P.G. Wolf. 2001. Metaxya lanosa, a second species in the genus and fern family Metaxyaceae. Systematic Botany 26:480-486.  

Wolf, P. G.,  D. R. Campbell, N. M. Waser, S. D. Sipes, T. R. Toler, and J. K. Archibald. 2001.  Tests of pre-and post-pollination barriers to hybridization between sympatric species of Ipomopsis (Polemoniaceae). American Journal of Botany 88: 213-219.

Barkworth, M.E.  2000.  Changing perceptions of the Triticeae.  Pp. 110-120 in S.W.L. Jacobs and J. Everett (Editors), Proceedings of the Third International Symposium on Grass Systematics and Evolution. CSIRO, Canberra, Australia.

Barkworth, M.E. and K.M. Capels.  2000.  The Poaceae in North America: A Geographic Perspective. Pp. 331-350 in S.W.L. Jacobs and J. Everett (Editors), Proceedings of the Third International Symposium on Grass Systematics and Evolution. CSIRO, Canberra, Australia.

Barkworth, M.E., K.M. Capels. and L.A. Vorobik. 2000. A grass manual for North America. Proc. Second Eastern Native Grass Symposium. p. 90.

Carman, J.G.  2000.  The gene effect: genome collisions and apomixis.  In:  Y Savidan and JG Carman, eds:  Advances in Apomixis Research.  FAO, CIMMYT, ORSTOM, Rome (in press).

Carman J.G.  2000.  The evolution of gametophytic apomixis.  In: T Batygina ed:  Embryology of Flowering Plants, Vol. 3: The Systems of Reproduction.  Russian Academy of Sciences, St. Petersburg (in press).

Ertter, B. 2000. Floristic surprises in North America north of Mexico. Annals of the Missouri Botanical Garden 87:81-109.

Gardner, D.R. and J.A. Pfister. 2000. Late season toxic alkaloid concentrations in tall larkspur (Delphinium spp.). J. Range Manage. 53:331-336.

Hinds, H. R.  2000. Flora of New Brunswick. Ed. 2.  Biology Department, University of New Brunswick, Fredericton, N.B., Canada

Jacobs, S.W.L., J. Everett, M.E. Barkworth, and C. Hsiao. 2000. Relationships within the Stipoid grasses (Gramineae). Pp. 75-82 in S.W.L. Jacobs and J. Everett (Editors), Proceedings of the Third International Symposium on Grass Systematics and Evolution. CSIRO, Canberra, Australia.  

Kelly, A.M. 2000. Guide to common macrolichens and bryophytes of the Umatilla National Forest. Umatilla National Forest, Pendleton, Oregon.  66 pp.

Larson, S.R., T.A. Jones, C. L. McCracken, Z-M Hu, and A.L. Palazzo.  2000.  Genetic diversity of bluebunch wheatgrass cultivars and a multiple-origin polycross.  Crop Science 40:  1142-1147.

Peters, R. May 1, 2000. Herbarium a well-kept secret.  The Herald Journal 11.

Pritchett, D. and F.J. Smith. 2000. 1999 Vegetation sampling at Nellis Air Force Bombing and Gunnery Range, Clark, Lincoln, and Nye counties, Nevada. Report to The Nature Conservancy, Nevada.

Smith, F.J. and D. Pritchett. 2000. Rare plant survey for Asclepias eastwoodiana at Nellis Air Force Bombing and Gunnery Range, Clark, Lincoln, and Nye counties, Nevada. Report to The Nature Conservancy, Nevada.

Valdes Reyna, J. and M.E. Barkworth. 2000.  Piptochaetium. Pp. 339-352 in Las Gramíneas de México, vol. 5. Secretario de Agricultura, Ganaderia y Desarrollo Rural, México, D.F., México.

Dobelis, P., J.E. Madl, J.A. Pfister, G.D. Manners, and J.P. Walrond. 1999. Effects of Delphinium alkaloids on neuromuscular transmission. J. Pharm. Exp. Therap.291:538-546.

Gardner, D.R., K.E. Panter, J.A. Pfister, and A.P. Knight. 1999. Analysis of toxic norditerpenoid alkaloids from Delphinium species by electrospray, atmospheric pressure chemical ionization and sequential tandem mass spectrometry. J. Agric. Food Chem. 47:5049-5058.

Kelly, A.M. 1999. Predictively mapping the plant associations of the North Fork John Day Wilderness in northeastern Oregon using classification tree modeling.  M.S. thesis, Department of Biology, Utah State University.

Panter, K.E., L.F. James, and D.R. Gardner. 1999.  Lupines, Poison-hemlock, and Nicotiana spp.: Toxicity and teratogenicity in livestock. J. Natural Toxins 891):117-134.

Pierson, K.1999. The Reproductive Biology and Edaphic Characteristics of a Rare, Gynodioecious Saxifrage: Saxifraga bryophora var. tobiasiae. Masters Thesis, Department of Biology, Utah State University. 154 pp.  

Pierson, K.  1999. Conservation Assessment and Conservation Strategy for Tobias’ Saxifrage, Saxifraga bryophora var. tobiasiae. Report to the  USFS Regional Office, Ogden, Utah and Payette National Forest, Supervisor’s Office, McCall, Idaho. 54 pp.

Pfister, J.A., D.R. Gardner, K.E. Panter, M.H. Ralphs, G.D. Manners, B.L. Stegelmeier, and T.K. Schoch. 1999. Larkspur (Delphinium spp.) toxicity to livestock. J. Nat. Toxins 8:81-94.

Pfister, J.A. and D.R. Gardner. 1999. Consumption of low larkspur (Delphinium nuttallianum) by cattle. J. Range Manage. 52:378-383.

Pierson, K. 1999. The reproductive biology and edaphic characteristics of a rare, gynodioecius saxifrage: Saxifraga bryophora var. tobiasiae. M.S. thesis, Department of Biology, Utah State University.

Shultz, L. M.  1999.  Using Geographic Information Systems in Floristic Studies.  Sida (in press).

Smith, F.J. 1999. An overlooked Phacelia from southern Nevada.  Submitted to The Nature Conservancy, Nevada. 

Weiss, R. 1999. Distribution of Pinus monophylla in the northern Wasatch Range of Utah. Great Basin Naturalist 59:292-294.

Wolf, P. G., S. D. Sipes, M. R. White, M. L. Martines, K. M. Pryer, A. R. Smith, K. Uedo. 1999. Phylogenetic relationships of the enigmatic fern families Hymenophyllopsidaceae and Lophosoriaceae: evidence from rbcL nucleotide sequences. Plant Systematics and Evolution: 219: 263-270.

Manners, G.D., K.E. Panter, J.A. Pfister, M.H. Ralphs, and L.F. James. 1998. The characterization and structure-activity evaluation of toxic norditerpenoid alkaloids from two Delphinium species. J. Nat. Prod. 61:1086-1089.

Panter, K.E., D.R. Gardner, and R.J. Molyneux. 1998. Teratogenic and fetotoxic effects of two piperidine alkaloid-containing lupines (L. formosus and L. arbustus) in cows. J. Natural Toxins 7(2):131-140.

Panter, K.E., D.R. Gardner, R.E. Shea, R.J. Molyneux and L.F. James. 1998. Toxic and teratogenic piperidine alkaloids from Lupinus, Conium, and Nicotiana species. pp. 345-350 in  T. Garland and A. C. Barr (Eds), Toxic Plants and other Natural Toxicants. CAB International, Wallingford, Oxon OX 108DE, united Kingdom.

Shultz, L. M., N. R. Morin & R. D. Ramsey.  1998.  Floristics in North America:  Tracking Rare Species Electronically in C. I. Peng, ed.  Rare, Threatened, and Endangered Floras of Asia and the Pacific Rim.  Institute of Botany, Academia Sinica, Monograph Series No. 16: 259--273.

Shultz, L. M..  1998.  A new species of Eriogonum (Polygonaceae: Eriogonoideae) from Utah and Nevada.  Harvard Papers in Botany 3(1): 49--52.

Shultz, L. M.  1998.  Flora of the Colorado Plateaus:  what do we know? In  L. M. Hill, ed. Learning from the Land:  Grand Staircase-Escalante National Monument Science Symposium Proceedings, pp. 203--209.   U. S. Dept. Interior, BLM/UT/GI-98/006+1220. 

Utah Division of Wildlife Resources.  1998.  Inventory of sensitive species and rare plants of Utah: an overview of their distribution and status.  Report to the Utah Reclamation Mitigation and Conservation Commission and the U.S. Department of the Interior.

West, N.E., R.J. Tausch, and P.T. Tueller. 1998. A management-oriented classification of Pinyon-Juniper woodlands of the Great Basin.  General Technical Report RMRS-GTR-12, U.S. Forest Service, U.S.D.A.

Barkworth, M.E.  1997.  Taxonomic and nomenclatural comments on the Triticeae in North America.  Phytologia 83:302-311. [Despite the date on the journal, this article was published in 1998].

Barkworth, M.E.  1997.  Elymus stebbinsii: Taxonomy, nomenclature, and distribution.  Phytologia 83:345-365. [Despite the date on the journal, this article was published in 1998].

Carlson, J.R. and M.E. Barkworth.  1997.  Elymus wawawaiensis: a species hitherto confused with Pseudoroegneria spicata (Triticeae, Poaceae).  Phytologia 83:312-330. [Despite the date on the journal, this article was published in 1998].  

Carman, J.G.  1997.  Asynchronous expression of duplicate genes in angiosperms may cause apomixis, bispory, tetraspory, and polyembryony.  Biol. J. Linn. Soc. 61:51-94.

Cronquist, A., N.H. Holmgren, and P.K. Holmgren.  1997.  Intermountain Flora, vol. 3a.  New York Botanical Garden, Bronx, New York.  

Gardner, D.R., G.D. Manners, M.H. Ralphs, and J.A. Pfister. 1997. Quantitative analysis of diterpenoid alkaloids in larkspur (Delphinium spp.) by Fourier Transform Infared Spectroscopy. Phytochem. Anal. 8:55-62.

Hou, Q., J.G. Carman, and W. A. Varga.  1997.  Micropropagation of sego lily (Calochortus nuttallii). Plant Cell Tis. Org. Cult. 49:149-151.

Pfister, J.A., D.R. Gardner, and K.W. Price. 1997. Grazing risk on tall larkspur-infested ranges. Rangelands 19(4):12-15.

Pfister, J.A., F.D. Provenza, G.D. Manners, D.R. Gardner, and M.H. Ralphs. 1997. Tall larkspur ingestion: can cattle regulate intake below toxic levels? J. Chem. Ecol. 23:759-777.

Pfister, J.A., M.H. Ralphs, G.D. Manners, D.R. Gardner, K.W. Price, and L.F. James. 1997. Early season grazing of tall larkspur- (Delphinium spp.) infested rangelands. J. Range Manage. 50: 391-398.

Ralphs, M.H., G.D. Manners, J.A. Pfister, D.R. Gardner, and L.F. James. 1997. Toxic alkaloid concentration in tall larkspur species in the western U.S. J. Range Manage. 50: 497-502.

Kropp, B.R., D.R. Hansen, P.G. Wolf, K.M Flint and S.V. Thomson. 1997. A study on the phylogeny of the dyer's woad rust fungus and other species of Puccinia from Crucifers. Phytopathology 87: 565-571. 

Panter, K.E., D.C. Baker, and P.O. Kechele, 1996. Water hemlock (Cicuta douglasii) toxicoses in sheep: pathological description and prevention of lesions and death. J. Vet. Diagn. Invest. 8:474-480.

Panter, K.E., D.R. Gay, C.C. Gay, L.F. James, R. Mills, J. M. Gaym and T.J. Baldwin. 1997. Observations of Lupinus sulphureus-induced "crooked calf disease". J. range Management 50:587-592.

Flora of North America Association.  1997. Cannabaceae, Ulmaceae, Urticaceae, and Moraceae.  Shultz, L. M., Taxon Editor, The Flora of North America, north of Mexico,  Vol. 3 : Magnoliidae and Hamamelidae.  Oxford University Press, New York.  (Shultz, L. M. as author of Trema Loureiro and Sherman-Broyles, S., Wm. Barker, and L.M. Shultz as authors of Ulmaceae)
Peel, M.D., J.G. Carman, and O. Leblanc.  1997.  Megasporocyte callose in apomictic buffelgrass, Kentucky bluegrass, Pennisetum squamulatum Fresen, Tripsacum L. and weeping lovegrass.  Crop Sci.  37:724-732.

Peel, M.D., J.G. Carman, Z-W. Liu, and R. R.-C. Wang.  1997.  Meiotic anomalies in hybrids between wheat and apomictic Elymus rectisetus (Nees in Lehm.) A. Löve & Connor.  Crop Sci.  37:717-723.

Sanders, Donald L. and Curtis Clark. 1987. Comparative morphology of the capitulum of Enceliopsis. Amer. J. Bot. 74:1072-1086.

Sipes, S. D. and P. G. Wolf. 1997. Clonal structure and patterns of allozyme diversity in the rare endemic Cycladenia humilis var. jonesii (Apocynaceae). American Journal of Botany 84: 401-409.

Smith, F.J. 1997. An inventory of rare, threatened, endangered, and endemic plants and unique communities on Nellis Air Force Bombing and Gunnery Range, Clark, Lincoln, and Nye Counties, Nevada.  Report to the U.S. Department of Defense, Department of the Air Force, produced in partnership with The Nature Conservancy of Nevada.  . 

Wolf, P. G., R. A. Murray, and S. D. Sipes. 1997. Species-independent, geographic structuring of chloroplast DNA haplotypes in a montane herb, Ipomopsis (Polemoniaceae). Molecular Ecology 6: 283-291.

Wolf, P. G. and R. B. Sinclair. 1997. Highly differentiated populations of the narrow endemic, Maquire primrose (Primula maguirei). Conservation Biology 11: 375-381

Wolf, P. G. 1997. Evaluation of atpB nucleotide sequences for phylogenetic studies of ferns and other pteridophytes. American Journal of Botany, 84: 1429-1440.

Barkworth, M.E., R.L. Burnham, and L.E. Talbert.  1996.  Elymus calderi: a new species in the Triticeae (Poaceae).  Systematic Botany 21:349-354.

Kropp, B.R. and L.J. Hutchison. 1996. Gigasperma americanum, a new hypogeous member of the Basidiomycota associated with Cercocarpus ledifolius in Utah. Mycologia 88: 662-665. 

Manners, G.D., R.Y. Wong, M. Benson, M.H. Ralphs, and J.A. Pfister. 1996. The characterization and absolute stereochemistry of barbaline, a diterpenoid alkaloid from Delphinium barbeyi. Phytochem. 42:875-879.

Manners, G.D., J.A. Pfister, and M.H. Ralphs. 1996. Larkspur: a poisonous problem on the range. Chemtech 26:49-54.

Manners, G.D., and J.A. Pfister. 1996. Sampling a poisonous plant population: quantifying toxic alkaloids in tall larkspur (Delphinium barbeyi) leaves. Weed Sci. 44:782-788.

Pierson, K. 1996. Permanent Monitoring Program for Happlopappus radiatus for Payette National Forest. Report to the  Idaho Conservation Data Center, Boise, ID and Payette National Forest, Supervisor’s Office, McCall, Idaho. 13 pp.    

Pierson, K. 1996. The Reproductive Biology and Edaphic Characteristic of a Rare Saxifrage: Saxifraga bryophora var. tobiasiae.  Report to the  Idaho Conservation Data Center, Boise, ID and Payette National Forest, Supervisor’s Office, McCall, Idaho. 14 pp.

Pfister, J.A., G.D. Manners, D.R. Gardner, K.W. Price, and M.H. Ralphs. 1996. Influence of alkaloid concentration on acceptability of tall larkspur (Delphinium spp.) to cattle and sheep. J. Chem. Ecol. 22:1147-1168.

Web Pages and Databases

Intermountain Herbarium. The official herbarium Web site is at  This is modified at long and irregular intervals.

Manual of Grasses for North America. The posted treatments for the Manual include distribution maps that are based on data compiled from many sources, including specimens in UTC.  These pages are also modified irregularly as no funding hase been provided for their development. 

Natural Heritage Program of Utah. The Intermountain Herbarium regularly provides the Natural Heritage Program with information. The herbarium benefits from the careful review of its specimens provided by 'Ben' Franklin, the botanist with the Natural Heritage Program.

Mindy Pratt, J. Bowns, R. Banner, A. Rasmussen. 2002. Range Plants of Utah.

Salo, C.  Earliest records of Red Brome.