Genomic Constitution of Taxa in Triticeae

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The table below presents genomic information on specimens in all the genera EXCEPT Triticum, Aegilops, and Amblyopryum which is on another page. The symbols used follow the recommendations of Wang et al. (1994). Notice that these authors recommended using E, not J, for the genome in T. elongatum and its allies (Thinopyrum as used on this web site), in order be consistent with previous use by wheat breeders. For information on this page I am indebted to Richard Wang and Bjorn Salomon. The information on the Triticum-Aegilops-Amblyopyrum page came from Richard Wang and Michael van Slageren.

The table below shows the genomic constitution of individual taxa sorted by genomes within affiliated groups of genera. For further information about these records, click here. To move to groups, use the following links: St genome species, E genome species, N genome species. Because of the way in which the species are grouped, it is impossible to make this as easy to use as might be desirable. One day I shall split it into multiple tables, each table including all taxa known to have a particular genome. When this happens, species polygenomic species will appear in more than one table.

RECENT WORK suggests that some of the constitutions reported on this page are not corrrect. Whether this is because the taxonomy of the species involved is not well understood or because the methods being used differ is not clear.

The last column contains the generic name that should be used if adopting the principle of genomic classification. It is based on the recommendations of Yen et al. . 2005, with one modification - the use of Thinopyrum rather than Lophopyrum when the two genera are combined in order to conform to the requiremens of the International Code of Botanical Nomenclature. Among the the principles Yen et al. followed in making their recommendations was that the treatment of genera should reflect both phylogenetic relationships and convenience and that force of habit should be considered, whenever possible, to avoid chaos. They acknowledge that their system is incomplete and does not resolve all problems. Prof. Yen sent me (Nov. 17, 2005) a working synopsis of Roegneria, for which I thank him. That information is being incorporated into the page for Roegneria. This page is designed to show which taxa have been genomically characterized and where the information was published.

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Taxon

Genomes

Reference

Genomic genus, see Yen et al. . 2005
Agropyron P   Agropyron
Crithopsis K    
Dasypyrum breviaristatum VbVb Galasso et al. 1997; Ohta and Morishita 2001  
 
Dasypyrum villosum V Galasso et al. 1997  
Pseudoroegneria St   Pseudoroegneria
Pseudoroegneria strigosa subsp. aegilopoides St Yu et al. (2008)  
Pseudoroegneria gracillima St Yu et al. (2008)  
Pseudoroegneria stipifolia St Yu et al. (2008)  
Pseudoroegnera strigosa subsp. strigosa St Yu et al. (2008)  
Pseudoroegneria libanotica St    
Pseudoroegneria tauri = Douglasdewey wangii StP Wang et al. (1986) Douglasdeweya
Roegneria magnicaespes St + not (E,P,H,Y) Yu et al. 2009 [DOI 10.1111/j.1756-1051.2009.00611.x] ?
St Ding et al. 2004; Ding et al. 2005; Zhang et al. 1999 Pseudoroegneria
Roegneria alashanica or Pseudoroegneria alashanica St + not (E,P,H,Y) Yu et al. 2009 [DOI 10.1111/j.1756-1051.2009.00611.x] ?
St Zhang et al. 1999 ; Zhang et al. 2002; Ding et al. 2004 Pseudoroegneria
Roegneria elytrigioides St Lu 1994; Ding et al. 2004 Pseudoroegneria
Pseudoroegneria cognata St Dewey 1981 Pseudoroegneria
Elymus agropyroides StH Dewey 1970b. Elymus
Elymus alaskanus StH Oergaard & Anamthawat-Jonsson 2001 Elymus
Elymus albicans StH Dewey 1970a. Elymus
Elymus canadensis StH Dewey 1970; Mason-Gamer 2001 Elymus

Elymus caninus

Type of Elymus sect. Goulardia [sect. name from 1941; Cynapoa is an older sectional name]

StH Dewey 1970; Redinbaugh et al. 2000 Elymus
StH Dewey 1974; Redinbaugh et al. 2000; Oergaard & Anamthawat-Jonsson 2001.
Elymus dasystachys StH Dewey 1967 [Need to check this] Elymus
StH Dewey 1970; Redinbaugh et al. 2000;
Elymus elymoides StH Dewey 1964; Redinbaugh et al. 2000; Mason-Gamer 2001; Larson et al. 2003.
Elymus hystrix StH Mason-Gamer 2001; Mason-Gamer et al. 2002. Elymus
Elymus glaucus StH Dewey 1965; Mason-Gamer 2001; Mason-Gamer et al. 2002. Elymus
Mason-Gamer et al. 2002.
Elymus lanceolatus StH Dewey 1967 (as Agropyron dasystachyum); Mason-Gamer 2001; Mason-Gamer et al. 2002. Elymus
Elymus magellanicus StH Jensen 1993. Elymus
Elymus multisetus StH Larson et al. 2003.. Elymus
Elymus mutabilis StH Dewey 1979; Redinbaugh et al. 2000. Elymus
Elymus pringlei StH Dewey 1976. Elymus
Elymus scribneri StH Dewey 1967. Elymus
Elymus sibiricus StH Dewey 1974. Elymus
Elymus subsecundus StH Dewey 1966. Elymus
Elymus tilcarensis StH Dewey 1977. Elymus
Elymus trachycaulus StH Dewey 1977; Redinbaugh et al. 2000 Elymus
StH Mason-Gamer et al. 2002.
Elymus virginicus StH Mason-Gamer 2001; Mason-Gamer et al. 2002. Elymus
Elymus wawawaiensis StH Mason-Gamer 2001; Mason-Gamer et al. 2002. Elymus
Elymus hoffmannii StStH Jensen & Asay 1996.  
Elymus transhyrcanus StStH Dewey 1972; Redinbaugh et al. 2000.  
Elymus repens StStH Assadi et Runemark 1994; Vershinin et al.1994  
StStH Assadi et Runemark 1994; Vershinin et al.1994  
StStH Redinbaugh et al. 2000  
XXH Oergaard & Anamthawat-Jonsson 2001  
StH(V-E-Ta-X) Mason-Gamer et al. 2002.  
Elymus patagonicus StHH Redinbaugh et al. 2000.  

Elymus dahuricus

Type of Elymus sect. Turczaninovia [sect. epithet 1932]

StHY Redinbaugh et al. 2000. Campeiostachys
Elymus drobovii StHY Dewey 1980 Campeiostachys
Elymus himalayanus StHY Lu and von Bothmer 1992. Campeiostachys
Elymus kamoji StHY Zhang et al. 1999. Campeiostachys
Elymus nutans StHY Lu 1993. Campeiostachys
Elymus schrenkianus StHY Lu and von Bothmer 1992. Campeiostachys
Elymus tsukushiensis StHY Lu et von Bothmer 1990a; Redinbaugh et al. 2000. Campeiostachys
Elymus caesifolius StStY Svitashev et al. 1998  
Elymus altissimus StY Lu and Salomon1992; Lu and von Bothmer 1993a. Roegneria
Elymus anthosachnoides StY Lu and Salomon1992; Lu et von Bothmer 1993a. Roegneria
Elymus barbicallus StY Lu and Salomon1992. Roegneria
Elymus breviglumis StY Lu and Salomon1992. Roegneria
Elymus brevipes StY Lu et von Bothmer 1993a. Roegneria
Elymus cacuminus StY Lu and Salomon 1993b. Roegneria

Elymus caucasicus

Type of Elymus sect. Clinelymopsis (1934)

StY Jensen and Wang 1991; Lu and von Bothmer 1993b; Redinbaugh et al. 2000. Roegneria
Elymus caucasicus StY Jensen and Wang 1991; Lu et von Bothmer 1993b; Redinbaugh et al. 2000 Roegneria
Elymus ciliaris StY Zhou et al. 1999; Redinbaugh et al. 2000; Mason-Gamer et al. 2002. Roegneria
Elymus ciliaris StY Zhou et al. 1999; Redinbaugh et al. 2000 Roegneria
Elymus dolichaterum StY Lu et Salomon1992. Roegneria
Elymus fedtschenkoi StY Liu et Dewey 1983. Roegneria
Elymus gmelinii StY Jensen et Hatch 1989; Lu et Salomon1992. Roegneria
Elymus grandis StStg (STg considered homoelogous with Y) Yu et al. 2009 [DOI 10.1111/j.1756-1051.2009.00611.x]  
StY Zhang et al. 1998a; Svitashev et al. 1998. Roegneria
Elymus jacquemontii StY Lu 1993. Roegneria
Elymus longearistatus StY Jensen et Wang 1991; Lu et von Bothmer 1993b. Roegneria
Elymus nevskii StY Lu et Salomon1992; Redinbaugh et al. 2000 Roegneria
Elymus nipponicus StY Lu et Salomon1992. Roegneria
Elymus panormitanus StY Lu et Salomon1992; Redinbaugh et al. 2000 Roegneria
Elymus parviglumis StY Lu et von Bothmer 1990b; Lu et Salomon 1993a. Roegneria
Elymus pendulinus StY Jensen 1990a; Zhou et al. 1999; Redinbaugh et al. 2000 Roegneria
Elymus praeruptus StY Salomon 1993. Roegneria
Elymus pseudonutans StY Lu et von Bothmer 1990b; Lu et Salomon1992. Roegneria
Elymus retroflexus StY Lu et Salomon 1993b. Roegneria
Elymus shandongensis StY Lu et Salomon1992. Roegneria
Elymus sclerus StY Lu et Salomon1992. Roegneria
Elymus semicostatus StY Lu et Salomon 1991, 1993a; Redinbaugh et al. 2000; Roegneria
Elymus strictus StY Lu et Salomon1992; Redinbaugh et al. 2000 Roegneria
Elymus tenuispicus StY Zhou et al. 1999 Roegneria
Elymus tibeticus StY Lu et Salomon1992, 1993a. Roegneria
Elymus validus StY Lu et Salomon1992. Roegneria
Elymus varius StY Lu et Salomon1992. Roegneria
Elymus yangiae StY Lu et Salomon1992; Lu et von Bothmer 1993a. Roegneria
Elymus borianus StStY Svitashev et al. 1998.  
Elymus caesifolius StStY Svitashev et al. 1998.  
Elymus glaucissimus StStY Lu et al. 1993; Redinbaugh et al. 2000  
Elymus tschimganicus StStY Lu et al. 1993; Jensen et al. 1994; Redinbaugh et al. 2000  
Kengyilia StPY Yen et Yang 1990 Kengyilia
Elymus alatavicus StPY Jensen et al. 1986; Redinbaugh et al. 2000 Kengyilia
Elymus batalinii StPY Jensen et al. 1986; Redinbaugh et al. 2000 Kengyilia
Elymus gobicola StPY Zhang et al. 2000. Kengyilia
Elymus grandiglumis StPY Jensen 1990b; Redinbaugh et al. 2000 Kengyilia
Elymus hirsuta StPY Zhang et al. 1999. Kengyilia
Elymus kengii StPY Jensen 1990b; Redinbaugh et al. 2000 Kengyilia
Elymus kokonoricus StPY Yang et al. 1992; Jensen 1996. Kengyilia
Elymus laxiflora StPY Zhang et al. 1998b. Kengyilia
Elymus melantherus StPY Yang et al. 1992; Jensen 1996. Kengyilia
Elymus muticus StPY Zhang et al. 1999. Kengyilia
Elymus retusus StPY Yang et al. 1992. Kengyilia
Elymus rigidulus StPY Yang et al. 1992; Zhang et al. 1999. Kengyilia
Elymus stenachyrus StPY Yang et al. 1992. Kengyilia
Elymus thoroldianus StPY Yang et al. 1992; Jensen 1996. Kengyilia
Elymus zhaosuensis StPY Zhang et al. 2000. Kengyilia
Elymus apricus StYW Stewart et al. 2005. Anthosachne
Douglasdeweya deweyi (Voucher SAUTI StP Jensen et al. 1992; Yen et al. 2005. Douglasdeweya
Douglasdeweya wangii = Pseudoroegneria tauri
(Voucher SAUTI 9109; grown from PI 401328)
StP Wang et al. . 1986; Yen et al. 2005. Douglasdeweya

Pseudoroegneria pertenuis
See note below

StP Wang et al. . 1986; Assadi 1995 but see Yen et al. 2005
Pascopyrum smithii StHNsXm Zhang and Dvorak 1991  
Elytrigia caespitosa subsp. caespitosa EeSt Yu et al. 2008 Thinopyrum
       
Pseudoroegneria geniculata subsp. scythica EeSt Liu et Wang 1992 Thinopyrum
Pseudoroegneria tauri
(Voucher PI 343188)
St Dewey 1974  
Elytrigia caespitosa subsp. caespitosa [combination in Thinopyrum is invalid] EeSt Liu et Wang 1989, 1993b Thinopyrum
Elytrigia caespitosa subsp. nodosa [combination not made] EeSt Liu et Wang 1993b Thinopyrum
Thinopyrum intermedium EbEeSt Liu et Wang 1993b; Xu et Conner 1994 Thinopyrum
EEsSt Chen et al. 1998
EeSt(V-Eb-R) Kishii et al. 2005.
Thinopyrum ponticum EEEStSt Zhang et al. 1996. Thinopyrum
EEEEE Muramatsu 1990; Wang et al. 1991.
EEEEsEs Chen et al. 1998.
Psammopyrum athericum EStP or ELP Ellneskog-Staam et al. 2001; .Ellneskog-Staam et al. 2003. Thinopyrum [combination not made]
Psammopyrum pungens EStStP or EStLP Ellneskog-Staam et al. 2003. Thinopyrum
Thinopyrum elongatum Ee Wang 1985 Thinopyrum
Thinopyrum bessarabicum Eb Wang 1985 Thinopyrum
Thinopyrum curvifolium EbEb Liu et Wang 1993a Thinopyrum
EcEcu McGuire 1984.
Thinopyrum distichum EbEe Liu et Wang 1993a Thinopyrum
Ed1Ed2 Pienaar et al. 1988; Liu et Wang 1993a.
Thinopyrum junceiforme EbEe Liu et Wang 1992; Refoufi et al. 2001 Thinopyrum
Thinopyrum junceum EbEbEe Liu et Wang 1993a Thinopyrum
Thinopyrum sartorii EbEe Liu et Wang 1992 Thinopyrum
Thinopyrum scirpeum EeEe Liu et Wang 1993a Thinopyrum
EsEsc McGuire 1984.
Thinopyrum pycnanthum = Elytrigia pycnantha EStP Refoufi et al. 2001 Thinopyrum
Crithopsis K    
Elymus rectisetus StYW Torabinejad et Mueller 1993; Redinbaugh et al. 2000 Anthosachne (in litt. 2005/10/30)
Elymus scaber StYW Torabinejad et Mueller 1993; Redinbaugh et al. 2000 Anthosachne (in litt. 2005/10/30)
Australopyrum W   Australopyrum
Elymus scabrifolius ?H Linde-Laursen et Seberg 2002  
Eremopyrum FXe Sakamoto 1979; Frederiksen and Bothmer 1995  
Festucopsis L    
Henrardia O    
Heteranthelium Q    
Hordeum bulbosum I    
Hordeum marinum Xa Bothmer et al. . 1986  
Hordeum murinum Xu Bothmer et al. . 1987, 1988a, 1998b  
Hordeum vulgare I    
Hordeum, other diploid species H    
Psathyrostachys Ns    
Hordelymus europaeus Ns? Svitashev et al. 1998.  
XoXr Bothmer et al. . 1994  
 
Ns(ancestor of St and L genomes) Petersen, G. and O. Seberg. 2008
Leymus aemulis NsXm Sun et al. . 1994.  
Leymus ambiguus NsXm Dewey 1976  
Leymus arenarius NsNsXmXm Wang et Jensen 1994.  
Leymus cinereus NsXm Wu et al. 2003.  
Leymus coreanus NsXm Jensen et Wang 1997.  
Elymus duthiei NsXm Svitashev et al. 1998  
Leymus erianthus NsXm Dubcovsky et al. 1997.  
Leymus flavescens NsXm Hole et al. 1999  
Elymus komarovii NsXm Svitashev et al. 1998  
Leymus mendocinus NsXm Dubcovsky et al. 1997.  
Leymus mollis NsXm Wang and Hsiao 1984.  
Leymus triticoides NsXm Wu et al. 2003.  
Leymus ugamicus NsXm Dewey 1980b.  
Leymus NsXm Zhang et Dvorak 1991; Wang et Jensen 1994  
Leymus californicus Ns? Jensen et Wang 1997; Dubcovsky et al. . 1997;  
Ns? Svitashev et al. 1998; Mason-Gamer 2001.  
NsXm Jensen et Wang 1997.  
Pascopyrum smithii StHNsXm Zhang and Dvorak 1991  
Peridictyon sanctus Xp Seberg et al. . 1991  
Australopyrum W Assigned by Löve 1984  
Elymus enysii HW Stewart et al. 2005 Stenostachys
Secale R    
Taeniatherum Ta    
Pseudoroegneria kosaninii not known    

Pseudoroegneria pertenuis: Assadi (1994, 1995) identified plants from Azerbaijan and those grown from the US accessions PI 40328 and 401330 as Elymus pertenuis = Pseudoroegneria pertenuis. Yen et al. . (2005) concluded that plants grown from seed of the two accessions were not the same as the type of pertenuis but belonged to a previously undescribed species, which they placed in their new genus Douglasdeweya as D. wangii. The accessions were originally identified as belonging to Pseudoreogneria tauri. According to Yen et al. . (2005), the genomic constitution of the pertenuis taxon is unknown.